But while a true four-rayed intersection, or simple cross, is
theoretically impossible (save as a transitory and highly unstable
condition), there is another condition which may closely simulate
it, and which is common enough. There are plenty of representations
of segmenting eggs, in which, instead of the triple junction and
polar furrow, the four cells (and in like manner their more numerous
successors) are represented as _rounded off_, and separated from one
another by an empty space, or by a little drop of an extraneous fluid,
evidently not directly miscible with the fluid surfaces of the cells.
Such is the case in the obviously accurate figure which Rauber gives
(Fig. 117, C) of the third mode of conjunction in the four-celled stage
of the frog’s egg. Here Rauber is most careful to point out that the
furrows do not simply “cross,” or meet in a point, but are separated
by a little space, which he calls the _Polgrübchen_, and asserts to be
constantly present whensoever the polar furrow, or _Brechungslinie_, is
not to be discerned. This little interposed space, with its contained
drop of fluid, materially alters the case, and implies a new condition
of theoretical and actual equilibrium. For, on the one hand, we see
that now the four intercellular partitions do not meet _one another
at all_; but really impinge upon four new and separate partitions,
which constitute interfacial contacts, not between cell and cell, but
between the respective cells and the intercalated drop. And secondly,
the angles at which these four little surfaces will meet the four
cell-partitions, will be determined, in the usual way, by the balance
between the respective tensions of these several surfaces. In an
extreme case (as in some pollen-grains) it may be found that the cells
under the observed circumstances are not truly in surface contact:
that they are so many drops which touch but do not “wet” one another,
and which are merely held together by the pressure of the surrounding
envelope. But even supposing, {313} as is in all probability the
actual case, that they are in actual fluid contact, the case from the
point of view of surface tension presents no difficulty. In the case of
the conjoined soap-bubbles, we were dealing with _similar_ contacts and
with _equal_ surface tensions throughout the system; but in the system
of protoplasmic cells which constitute the segmenting egg we must make
allowance for _an inequality_ of tensions, between the surfaces where
cell meets cell, and where on the other hand cell-surface is in contact
with the surrounding medium,—in this case generally water or one of the
fluids of the body. Remember that our general condition is that, in our
[Illustration: Fig. 118.]
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