Let us imagine a system of equal cylinders, or equal spheres, in
contact with one another in a plane, and represented in section by
the equal and contiguous circles of Fig. 121. I borrow my figure, by
the way, from an old Italian naturalist, Bonanni (a contemporary of
Borelli, of Hay and Willoughby and of Martin Lister), who dealt with
this matter in a book chiefly devoted to molluscan shells[362].
It is obvious, as a simple geometrical fact, that each of these equal
circles is in contact with six surrounding circles. Imagine now that
the whole system comes under some uniform stress. It may be of uniform
surface tension at the boundaries of all the cells; it may be of
pressure caused by uniform growth or expansion within the cells; or
it may be due to some uniformly applied constricting pressure from
without. In all of these cases the _points_ of contact between the
circles in the diagram will be extended into {319} _lines_ of contact,
representing _surfaces_ of contact in the actual spheres or cylinders;
and the equal circles of our diagram will be converted into regular and
equal hexagons. The angles of these hexagons, at each of which three
hexagons meet, are of course angles of 120°. So far as the form is
concerned, so long as we are concerned only with a morphological result
and not with a physiological process, the result is precisely the same
whatever be the force which brings the bodies together in symmetrical
apposition; it is by no means necessary for us, in the first instance,
even to enquire whether it be surface tension or mechanical pressure or
some other physical force which is the cause, or the main cause, of the
phenomenon.
[Illustration: Fig. 121. Diagram of hexagonal cells. (After Bonanni.)]
The production by mutual interaction of polyhedral cells, which, under
conditions of perfect symmetry, become regular hexagons, is very
beautifully illustrated by Prof. Bénard’s “_tourbillons cellulaires_”
(cf. p. 259), and also in some of Leduc’s diffusion experiments. A weak
(5 per cent.) solution of gelatine is allowed to set on a plate of
glass, and little drops of a 5 or 10 per cent. solution of ferrocyanide
of potassium are then placed at regular intervals upon the gelatine.
Immediately each little drop becomes the centre, or pole, of a system
of diffusion currents, {320} and the several systems conflict with and
repel one another, so that presently each little area becomes the seat
of a double current system, from its centre outwards and back again;
until at length the concentration of the field becomes equalised and
the currents {321}
[Illustration: Fig. 122. An “artificial tissue,” formed by coloured
drops of sodium chloride solution diffusing in a less dense solution of
the same salt. (After Leduc.)]
[Illustration: Fig. 123. An artificial cellular tissue, formed by the
diffusion in gelatine of drops of a solution of potassium ferrocyanide.
(After Leduc.)]
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