What has happened here is not difficult to understand. Imagine, as
before, a system of equal spheres all in contact, each one therefore
touching six others in an equatorial plane; and let the cells be not
only in contact, but become attached at the points of contact. Then
instead of each cell expanding, so as to encroach on and fill up the
intercellular spaces, let each cell tend to contract or shrivel up,
by the withdrawal of fluid from its interior. The {336} result will
obviously be that the intercellular spaces will increase; the six
equatorial attachments of each cell (Fig. 133, _a_) (or its twelve
attachments in all, to adjacent cells) will remain fixed, and the
portions of cell-wall between these points of attachment will be
withdrawn in a symmetrical fashion (_b_) towards the centre. As the
final result (_c_) we shall have a “dodecahedral star” or star-polygon,
which appears in section as a six-rayed figure. It is obviously
necessary that the pith-cells should not only be attached to one
another, but that the outermost layer should be firmly attached to
a boundary wall, so as to preserve the symmetry of the system. What
actually occurs in the rush is tantamount to this, but not absolutely
identical. Here it is not so much the pith-cells which tend to shrivel
within a boundary of constant size, but rather the boundary wall (that
is, the peripheral ring of woody and other tissues) which continues to
expand after the pith-cells which it encloses have ceased to grow or
to multiply. The twelve points of attachment on the spherical surface
of each little pith-cell are uniformly drawn asunder; but the content,
or volume, of the cell does not increase correspondingly; and the
remaining portions of the surface, accordingly, shrink inwards and
gradually constitute the complicated surface of a twelve-pointed star,
which is still a symmetrical figure and is still also a surface of
minimal area under the new conditions.
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Public-domain text, read in full here on John Shaqi.
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