While in very many cases the partitions (like the walls of the original
cell) will be either plane or spherical, a more complex curvature will
be assumed under a variety of conditions. It will be apt to occur,
for instance, when the mother-cell is irregular in shape, and one
particular case of such asymmetry will be that in which (as in Fig.
143) the cell has begun to branch, or give off a diverticulum, before
division takes place. A very complicated case of a different kind,
though not without its analogies to the cases we are considering, will
occur in the partitions of minimal area which subdivide the spiral
tube of a nautilus, as we shall presently see. And again, whenever we
have a marked internal asymmetry of the cell, leading to irregular
and anomalous modes of division, in which the cell is not necessarily
divided into two equal halves and in which the partition-wall may
assume an oblique position, then apparently anomalous curvatures will
tend to make their appearance[388].
Suppose that a more or less oblong cell have a tendency to divide by
means of an oblique partition (as may happen through various causes
or conditions of asymmetry), such a partition will still have a
tendency to set itself at right angles to the rigid walls {356} of the
mother-cell: and it will at once follow that our oblique partition,
throughout its whole extent, will assume the form of a complex,
saddle-shaped or anticlastic surface.
[Illustration: Fig. 143. Diagrammatic explanation of S-shaped
partition.]
Many such cases of partitions with complex or double curvature exist,
but they are not always easy of recognition, nor is the particular
case where they appear in a _terminal_ cell a common one. We may see
them, for instance, in the roots (or rhizoids) of Mosses, especially
at the point of development of a new rootlet (Fig. 142, C); and again
among Mosses, in the “paraphyses” of the male prothalli (e.g. in
_Polytrichum_), we find more or less similar partitions (D). They are
frequent also among many Fuci, as in the hairs or paraphyses of Fucus
itself (B). In _Taonia atomaria_, as figured in Reinke’s memoir on
the Dictyotaceae of the Gulf of Naples[389], we see, in like manner,
_oblique_ partitions, which on more careful examination are seen to be
curves of double curvature (Fig. 142, A).
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