Examples of these various arrangements meet us at every turn, and
not only in cell-aggregates, but in various cases where non-rigid
and semi-fluid partitions (or partitions that were so to begin
with) meet together. And it is a necessary consequence of this
physical phenomenon, and of the limited and very small number of
possible arrangements, that we get similar appearances, capable of
representation by the same diagram, in the most diverse fields of
biology[393].
[Illustration: Fig. 159.]
Among the published figures of embryonic stages and other cell
aggregates, we only discern these little intermediate partitions in
cases where the investigator has drawn carefully just what lay before
him, without any preconceived notions as to radial or other symmetry;
but even in other cases we can generally recognise, without much
difficulty, what the actual arrangement was whereby the cell-walls
met together in equilibrium. I have a strong suspicion that a leaning
towards Sachs’s Rule, that one cell-wall tends to set itself at right
angles to another cell-wall (a rule whose strict limitations, and
narrow range of application, we have already {377} considered) is
responsible for many inaccurate or incomplete representations of the
mutual arrangement of aggregated cells.
[Illustration: Fig. 160. Segmenting egg of _Trochus_. (After Robert.)]
[Illustration: Fig. 161. Two views of segmenting egg of _Cynthia
partita_. (After Conklin.)]
[Illustration: Fig. 162. (_a_) Section of apical cone of _Salvinia_.
(After Pringsheim[394].) (_b_) Diagram of probable actual arrangement.]
[Illustration: Fig. 163. Egg of _Pyrosoma_. (After Korotneff).]
[Illustration: Fig. 164. Egg of _Echinus_, segmenting under pressure.
(After Driesch.)]
In the accompanying series of figures (Figs. 160–167) I have {378}
set forth a few aggregates of eight cells, mostly from drawings of
segmenting eggs. In some cases they shew clearly the manner in which
the cells meet one another, always at angles of 120°, and always with
the help of five intermediate boundary walls within the eight-celled
system; in other cases I have added a slightly altered drawing, so as
to shew, with as little change as {379} possible, the arrangement
of boundaries which probably actually existed, and gave rise to the
appearance which the observer drew. These drawings may be compared
with the various diagrams of Fig. 158, in which some seven out of the
possible thirteen arrangements of five intermediate partitions (for a
system of eight cells) have been already set forth.
[Illustration: Fig. 165. (_a_) Part of segmenting egg of Cephalopod
(after Watase); (_b_) probable actual arrangement.]
[Illustration: Fig. 166. (_a_) Egg of _Echinus_; (_b_) do. of _Nereis_,
under pressure. (After Driesch).]
[Illustration: Fig. 167. (_a_) Egg of frog, under pressure (after
Roux); (_b_) probable actual arrangement.]
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