The figures given by
Goebel of the development of the pollen of Neottia (3, _a_–_e_: all the
figures referring to grains taken from a single anther), illustrate
this to perfection; and it will be seen that, when the four cells lie
in a plane, they conform exactly to our typical diagram of the first
four cells in a segmenting ovum. Occasionally, though the four cells
lie in a plane, the diagram seems to fail us, for the cells appear to
meet in a simple cross (as in 5); but here we soon perceive that the
cells are not in complete interfacial contact, but are kept apart by a
little intervening drop of fluid or bubble of air. The spores of ferns
(7) develop in very much the same way as pollen-grains; and they also
very often retain traces of the shape which they assumed as members of
a tetrahedral figure. Among the “tetraspores” (8) of the Florideae, or
Red Seaweeds, we have a phenomenon which is in every respect analogous.
Here again it is obvious that, apart from differences in actual
magnitude, and apart from superficial or “accidental” differences
(referable to other physical phenomena) in the way of colour, {398}
texture and minute sculpture or pattern, it comes to pass, through the
laws of surface-tension and the principles of the geometry of position,
that a very small number of diagrammatic figures will sufficiently
represent the outward forms of all the tetraspores, four-celled
pollen-grains, and other four-celled aggregates which are known or are
even capable of existence.
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Public-domain text, read in full here on John Shaqi.
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