On the growing root of a bean, ten narrow zones were marked off,
starting from the apex, each zone a millimetre in breadth. After
twenty-four hours’ growth, at a certain constant temperature, the
whole marked portion had grown from 10 mm. to 33 mm. in length; but
the individual zones had grown at very unequal rates, as shewn in the
annexed table[129].
Zone Increment
mm.
Apex 1·5
2nd 5·8
3rd 8·2
4th 3·5
5th 1·6
6th 1·3
7th 0·5
8th 0·3
9th 0·2
10th 0·1
{96}
[Illustration: Fig. 20. Rate of growth in successive zones near the tip
of the bean-root.]
The several values in this table lie very nearly (as we see by Fig.
20) in a smooth curve; in other words a definite law, or principle of
continuity, connects the rates of growth at successive points along the
growing axis of the root. Moreover this curve, in its general features,
is singularly like those acceleration-curves which we have already
studied, in which we plotted the rate of growth against successive
intervals of time, as here we have plotted it against successive
spatial intervals of an actual growing structure. If we suppose for a
moment that the velocities of growth had been transverse to the axis,
instead of, as in this case, longitudinal and parallel with it, it is
obvious that these same velocities would have given us a leaf-shaped
structure, of which our curve in Fig. 20 (if drawn to a suitable scale)
would represent the actual outline on either side of the median axis;
or, again, if growth had been not confined to one plane but symmetrical
about the axis, we should have had a sort of turnip-shaped root, {97}
having the form of a surface of revolution generated by the same
curve. This then is a simple and not unimportant illustration of the
direct and easy passage from velocity to form.
A kindred problem occurs when, instead of “zones” artificially marked
out in a stem, we deal with the rates of growth in successive actual
“internodes”; and an interesting variation of this problem occurs when
we consider, not the actual growth of the internodes, but the varying
number of leaves which they successively produce. Where we have whorls
of leaves at each node, as in Equisetum and in many water-weeds, then
the problem presents itself in a simple form, and in one such case,
namely in Ceratophyllum, it has been carefully investigated by Mr
Raymond Pearl[130].
It is found that the mean number of leaves per whorl increases with
each successive whorl; but that the rate of increment diminishes from
whorl to whorl, as we ascend the axis. In other words, the increase
in the number of leaves per whorl follows a logarithmic ratio; and if
_y_ be the mean number of leaves per whorl, and _x_ the successional
number of the whorl from the root or main stem upwards, then
_y_ = _A_ + _C_ log(_x_ − _a_),
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