It is evident, then, that in this case of “dimorphism,” the tails of
the one group of earwigs (which Bateson calls the “high males”) have
either grown _faster_, or have been growing for a longer period of
time, than those of the “low males.” If we could be certain that the
whole random sample of earwigs were of one and the same age, then we
should have to refer the phenomenon of dimorphism to a physiological
phenomenon, simple in kind (however remarkable and unexpected); viz.
that there were two alternative {106} values, very different from
one another, for the mean velocity of growth, and that the individual
earwigs varied around one or other of these mean values, in each case
according to the law of probabilities. But on the other hand, if we
could believe that the two groups of earwigs were _of different ages_,
then the phenomenon would be simplicity itself, and there would be no
more to be said about it[133].
――――――――――
Before we pass from the subject of the relative rate of growth of
different parts or organs, we may take brief note of the fact that
various experiments have been made to determine whether the normal
ratios are maintained under altered circumstances of nutrition, and
especially in the case of partial starvation. For instance, it has been
found possible to keep young rats alive for many weeks on a diet such
as is just sufficient to maintain life without permitting any increase
of weight. The rat of three weeks old weighs about 25 gms., and under a
normal diet should weigh at ten weeks old about 150 gms., in the male,
or 115 gms. in the female; but the underfed rat is still kept at ten
weeks old to the weight of 25 gms. Under normal diet the proportions
of the body change very considerably between the ages of three and ten
weeks. For instance the tail gets relatively longer; and even when the
_total_ growth of the rat is prevented by underfeeding, the _form_
continues to alter so that this increasing length of the tail is still
manifest[134]. {107}
_Full-fed Rats._
Age in Length of Length of Total
weeks body (mm.) tail (mm.) length % of tail
0 48·7 16·9 65·6 25·8
1 64·5 29·4 93·9 31·3
3 90·4 59·1 149·5 39·5
6 128·0 110·0 238·0 46·2
10 173·0 150·0 323·0 46·4
_Underfed Rats._
6 98·0 72·3 170·3 42·5
10 99·6 83·9 183·5 45·7
Again as physiologists have long been aware, there is a marked
difference in the variation of weight of the different organs,
according to whether the animal’s total weight remain constant, or
be caused to diminish by actual starvation; and further striking
differences appear when the diet is not only scanty, but ill-balanced.
But these phenomena of abnormal growth, however interesting from
the physiological view, are of little practical importance to the
morphologist.
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