On Molecular and Microscopic Science, Volume 1 (of 2)Somerville, Mary
History
On Molecular and Microscopic Science, Volume 1 (of 2)
Somerville, Mary
Matter -- Constitution; Microscopy; Natural history
[Illustration: Fig. 8. Volvox globator.]
The Volvocineæ consist of various species according as the internal
matter of the primordial cell divides into 2, 4, 8, 32, or a greater
number of equal parts, forming respectively as many free cells which
ultimately become ciliated spores, by means of which the globe either
rotates on the spot, or in straight lines. The Volvox globator found in
fresh-water pools is one of the most remarkable of these, both for its
peculiarity and beauty of structure and for its comparatively large
size, since in some lights it is visible to the naked eye while swimming
in a drop of water. When viewed with a microscope, it is a pellucid
sphere whose surface is studded with green spots, often connected by
green threads; each of the spots has two cilia, so that the surface is
bristled with these filaments, whose vibrations give the sphere either a
rolling or smooth motion, or make it spin like a top in the same place.
In the interior of the sphere there are from two to twenty dark green
globes of different sizes; the smaller are attached to the internal
surface, while the larger rotate freely by their cilia in the internal
cavity. After a time the sphere bursts open and its inhabitants swim
forth, and soon assume the form and character of that which gave them
birth.
The growth and development of the Volvox globator are peculiar, for in
the primordial cell the red and green endochrome breaks up into numerous
angular masses, and a central globe rather larger than the rest. The
angular masses are connected by green threads, the interstices between
all the bodies are filled with a hyaline substance secreted from their
surfaces, and the whole is enclosed in a distinctly membranous globular
envelope.
As this young Volvox increases gradually in size, the hyaline matter is
increased, the green threads lengthen, and the angular masses assume the
form of a flask the 1/3000th of an inch in diameter exactly as in the
Protococcus; for the green matter with a few red spots is collected in
the thick end, while the hyaline beak is turned towards the
circumference of the sphere, which is pierced by their long cilia. Each
of them is invested with a pellucid envelope of considerable thickness,
the borders of which are flattened against those of similar envelopes.
While these ciliated bodies are approaching maturity their endochrome
exhibits vacuoles or apparently empty cavities of a spherical form about
one-third of its own diameter. Mr. G. Busk discovered that these
vacuoles expand and contract at regular intervals of about forty
seconds. The contraction, which almost obliterates the cavity of the
vacuole, is rapid and sudden; the dilatation is slow and gradual. This
action ceases when the body comes to maturity.
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