On Molecular and Microscopic Science, Volume 1 (of 2)Somerville, Mary
History
On Molecular and Microscopic Science, Volume 1 (of 2)
Somerville, Mary
Matter -- Constitution; Microscopy; Natural history
The systematic arrangement of the Ferns is chiefly founded on
peculiarities of the sori and sporangia, characters which are quite
intelligible by the aid of a good magnifying lens, these spore-cases
being very pretty opaque objects under the microscope. Thus some of the
primary divisions are founded on the presence or absence of the ring or
annulus on the spore-case. Another series of divisions are founded
mainly on the nature of the ring in those cases when it is present; and,
for the rest, the form and position of the sori come in as
discriminating characters. In this way the main groups are marked out,
but, in the case of the genera, still further recourse is had to the
sori and its covering, and by some modern authors supplementary
characters derived from the venation are brought into use. The following
is the arrangement adopted by Mr. Moore,[71] and which agrees in its
general features with that of most modern observers.
The two leading groups are the _Annulate_ and the _Exannulate_ Ferns,
the first being much the larger division, and consisting of the order
Polypodiaceæ, which comprises the True Ferns, while the second includes
the two orders Marattiaceæ and Ophioglossaceæ, which are some times
called Pseudo-Ferns.
The POLYPODIACEÆ, distinguished by the ring or annulus, which more or
less completely girts the sporangia, offer so much variety of
structure that it becomes necessary to subdivide them; and for this
purpose characters derived from the form, number, or position of the
sporangia, or the structure or development of the ring, are chiefly
relied on. This gives several groups, _e.g._ Polypodineæ, the most
comprehensive of all, including some ten or twelve minor groups, in
which the sporangia are almost equally convex, and have a vertical and
nearly complete ring, and in which the dehiscence is transverse at a
part called the stoma, where the striæ of the ring are elongated, and
apparently weaker; Cyatheineæ, in which the spore-cases are sessile or
nearly so, seated on an elevated receptacle, with the nearly complete
ring more or less obliquely vertical, that is, vertical below, curving
laterally towards the top, and the dehiscence transverse; Matonineæ, a
single species only, in which the sporangia are sessile, bursting
horizontally, not vertically, the ring being broad, suboblique, and
nearly complete, and the dorsal sori oligocarpous, covered by
umbonato-hemispherical indusia, affixed by a central stalk;
Gleichenineæ, with the ring complete, transverse, either truly or
obliquely horizontal, the sporangia globose-pyriform, forming
oligocarpous sori, _i.e._ sori consisting of but few spore-cases (two
or four to ten or twelve), situated at the back of the frond, sessile
or nearly so, and bursting vertically, the fronds, moreover, being
rigid and opaque, and usually dichotomously-branched; Trichomanineæ,
with the ring resembling that of the Gleichenineæ, but the sporangia
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