On Molecular and Microscopic Science, Volume 1 (of 2)Somerville, Mary
History
On Molecular and Microscopic Science, Volume 1 (of 2)
Somerville, Mary
Matter -- Constitution; Microscopy; Natural history
The fronds of the Bristle fern, Trichomanes radicans, are somewhat
deltoid, or of a more elongated form, flimsy, and beautifully
reticulated when viewed with a microscope. They rise at intervals from a
creeping rhizome, bristly, with narrow articulate scales, which often
covers the most precipitous rocks on which it grows with a dark-coloured
network. The rachis is branched and rebranched three or four times, and
the whole being distinctly winged on both sides in the plane of the
frond, the successive branches and branchlets running through the leafy
part become the veins of those parts, so that a segment of the frond is
merely a winged vein, the wings on the branches being however broader
than those on the rachis. The veins, which divide alternately, are hard,
woody, and wire-like, and, when barren, terminate before reaching the
ends of the segments; but, when fertile, they extend beyond the segment,
the tissue of which separates and distends in the form of a more or less
elongated cup around the prolonged vein. At the base of that vein the
sporangia form a small globular cluster; and, as they advance towards
maturity, the vein extends in the form of a bristle, far beyond the
mouth of the cup (fig. 68). The cup is winged in the more luxuriant form
of the plant, in consequence of the double layer of tissue composing the
segment to which it belongs not separating through its whole breadth.
The cup-like cylinder being slightly compressed in the plane of the
frond is indicative of its origin.
[Illustration: Fig. 69. Hymenophyllum tunbridgense.]
A slight difference in fructification separates the Hymenophyllum
tunbridgense, or Film fern (fig. 69), from the Trichomanes, for both
have a creeping rhizome, from which single fronds spring at short
intervals. The leafy parts of the fronds are merely winged veins, and
the wings consist of two layers of cellular tissue finely reticulated.
The Hymenophyllum is distinguished from the Trichomanes by a two-valved
cup spinously serrated, which completely conceals the fruit-bearing
vein, while, in the Trichomanes, the cup is more cylindrical,
smooth-edged, with the fertile vein projecting far beyond it (fig. 68).
The beautiful little Film fern, with its semi-transparent fronds not
more than two or three inches high, grows on shady wet rocks, among the
moss on the branches and roots of old trees, and on the ground near
lakes and rivulets. It is hardy enough to live in the Highlands of
Scotland; but the H. unilaterale, a far less beautiful plant, has its
limit in Unst, the most northern of the Shetland Islands. The
fructification in the latter plant, though the same in position, is
stalked instead of being sessile. The involucre is rounded and ovate,
instead of being cup-shaped, with much swollen convex valves, meeting by
their edges but not compressed towards the apex.
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