On Molecular and Microscopic Science, Volume 1 (of 2)Somerville, Mary
History
On Molecular and Microscopic Science, Volume 1 (of 2)
Somerville, Mary
Matter -- Constitution; Microscopy; Natural history
In all common Orchids there is only one stamen, which is confluent with
the three pistils, to form what is called the column. The edges of the
labellum are attached to the sides of the column, leaving a space known
as the chamber or mouth of the nectary, and thus the mouth has the
column on one side, and the labellum on the other. Although the three
pistils are united into one, the three stigmas are not; for the stigma
of the upper pistil is transformed into a pouch-shaped viscous mass
(_r_, fig. 73), called the rostellum, bearing no resemblance whatever to
a stigma, while the other two stigmas form a bi-lobed confluent mass or
stigma below it, through which the pollen grains fertilize the ovary,
which latter forms the apparently twisted stalk, by which the blossom is
attached to the stem. In fact, the rostellum (_r_) projects into the
mouth of the nectary, and overhangs the confluent stigmas (_s s_). The
lowest and narrowest part of a hood-shaped anther (fig. 73 _a a_,) is
attached to the back of the rostellum, and consists of two rather widely
separated oblong cells, which open longitudinally in front. Each cell
contains a pollen-mass, or pollinium. Fig. 74 represents one with its
club-shaped mass of pollen grains, its stalk, and viscid disc. These
objects are seen in situ in fig. 73. A pollinium consists of a number of
wedge-shaped packets of pollen grains held together by exceedingly
elastic fine threads, as in fig. 75; the packets unite into the
club-shaped head, and the threads form the stem; the viscid discs are
formed by the rostellum.
[Illustration: Fig. 74. Pollinium of Orchis mascula:—_p_, pollinium;
_c_, caudicle; _d_, disc.]
[Illustration: Fig. 75. Pollen grains of Orchis mascula.]
At an early period of growth the rostellum consists of a mass of
polygonal cells full of brownish matter, which soon resolve themselves
into two balls of an extremely viscid semifluid substance void of
structure. These two discs are flat on the top, and rounded below. They
lie quite free within the rostellum, except at the back, where each
viscid disc firmly adheres to a small portion of the exterior membrane
of the rostellum. The ends of the two stalks of the pollinia, or
granular masses, are strongly attached to these little discs, or balls.
[Illustration: Fig. 76. Pollinia of Orchis mascula, showing front view
of the discs and caudicles within the rostellum:—_r_, rostellum; _c_,
caudicle; _d_, disc.]
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