On Molecular and Microscopic Science, Volume 2 (of 2)Somerville, Mary
History
On Molecular and Microscopic Science, Volume 2 (of 2)
Somerville, Mary
Matter -- Constitution; Microscopy; Natural history
The spines of the Echinus miliaris, of which fig. 141 represents the
segment of a section highly magnified, are fluted columns of calcareous
glass, the grooves of which are filled with solid glassy matter curved
on the exterior. The innumerable hair-like objects attached to the
shells of some of the Echinidæ, the almost filamental spines of others,
and the pedicellariæ themselves, are formed of a regularly reticulated
substance. When the Echinidæ are stripped of their spines and all their
appendages, their shells show 2,400 plates united with the symmetry of a
tesselated pavement.
[Illustration: Fig. 142. Pluteus of the Echinus:—_a_, mouth; _b_,
stomach; _c_, echinoid disk; _d d d d_, four arms of the pluteus-body;
_e_, calcareous framework; _f_, ciliated lobes; _g g g g_, ciliated
processes of the proboscis.]
The Echinidæ are male and female, and the eggs are excluded through the
five perforated productive plates at the posterior end of the shell.
According to the observations of Prof. Fritz Müller the embryo, soon
after issuing from the egg, takes a form represented (magnified) in fig.
142.
All parts of this creature, which is called a Pluteus, are strengthened
by a framework of calcareous rods tipped with orange colour, all the
rest being transparent and colourless. It swims freely, back foremost,
by means of its cilia.
[Illustration: Fig. 143. Larvæ of Echinus in various stages of
development within the Pluteus, which is not represented:—B, disk with
the first indication of the cirrhi; C, disk with the origin of the
spines between the cirrhi; D, more advanced disk with the cirrhi, _g_,
and spines, _x_, projecting from the surface.]
While in this active state a circular disk (_c_, fig. 142), covering the
stomach (_b_, fig. 142), appears within it, which gradually expands, and
sends through the skin of the Pluteus spines, pedicellariæ, and
tubercules, ultimately developed into hollow feet. Then the feet are
pushed out and drawn in, the pedicellariæ (D, fig. 143) snap their
pincers; and while the half-formed Echinus is making these motions
within the Pluteus, the mouth and gullet of the Pluteus itself are in
constant activity; and, while it swims about, the unformed Echinus
within it gets a globular shape, the shell is formed, and when the
Echinus is complete, the rest of the Pluteus is thrown off, and the
young animal rolls away.
The free swimming larval zooids of the Echinodermata are generally
hyaline, and some are phosphorescent. The Pluteus is also the larval
zooid of the ophiurid star-fishes; they may be seen in great numbers on
the surface of the sea in August and September. The young star-fish is
formed in them by a process analogous to that described. The motions of
the Echinidæ are reflex; nothing indicates volition.
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