On the Genesis of SpeciesMivart, St. George Jackson
Religion
On the Genesis of Species
Mivart, St. George Jackson
Evolution (Biology)
Again, Nicholas Wagner found in certain other Diptera, the Hessian flies,
that the larva gives rise to secondary larvae within it, which develop and
burst the body of the primary larva. The secondary larvae give rise,
similarly, to another set within them, and these again to another[171] set.
Again, the fact that in _Taenia echinococcus_ one egg produces numerous
individuals, tends to invalidate the argument that the increase of segments
during development is a relic of specific genesis.
Mr. H. Spencer seems to deny serial homology to the mollusca, but it is
difficult to see why the shell segments of chiton are not such homologues
because the segmentation is superficial. Similarly the external processes
of eolis, doris, &c., are good examples of serial homology, as also are
plainly the successive chambers of the orthoceratidae. Nor are parts of a
series less serial, because arranged spirally, as in most gasteropods. Mr.
Spencer observes of the molluscous as of the vertebrate animal, "You cannot
cut it into transverse slices, each of which contains a digestive organ, a
respiratory organ, a reproductive organ, &c."[172] But the same may be said
of every single arthropod and annelid if it be meant that all these organs
are not contained in every possible slice. While if it be meant that parts
of all such organs are contained in certain slices, then some of the
mollusca may also be included.
Another objection to Mr. Spencer's speculation is derived from
considerations which have already been stated, as to past time. For if{171}
the annulose animals have been formed by aggregation, we ought to find this
process much less perfect in the oldest form. But a complete development,
such as already obtains in the lobster, &c., was reached by the Eurypterida
and Trilobites of the palaeozoic strata; and annelids, probably formed
mainly like those of the present day, abounded during the deposition of the
oldest fossiliferous rocks.
[Illustration: TRILOBITE.]
Thirdly, and lastly, as regards such serial homology as is exemplified by
the backbone of man, there are also several objections to Mr. Spencer's
mechanical explanation.
On the theory of evolution most in favour, the first Vertebrata were
aquatic. Now, as natation is generally effected by repeated and vigorous
lateral flexions of the body, we ought to find the segmentation much more
complete laterally than on the dorsal and ventral aspects of the spinal
column. Nevertheless, in those species which, taken together, constitute a
series of more and more distinctly segmented forms, the segmentation
gradually increases _all round_ the central part of the spinal column.
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