On the Genesis of SpeciesMivart, St. George Jackson
Religion
On the Genesis of Species
Mivart, St. George Jackson
Evolution (Biology)
Professor Burt G. Wilder, in his paper on extra digits,[188] has {181}
recorded no less than twenty-four cases where such excess coexisted in both
little fingers; also one case in which the right little finger and little
toe were so affected; six in which it was both the little fingers and both
the little toes; and twenty-two other cases more or less the same, but in
which the details were not accurately to be obtained.
Mr. Darwin cites[189] a remarkable instance of what he is inclined to
regard as the development in the foot of birds of a sort of representation
of the wing-feathers of the hand. He says: "In several distinct breeds of
the pigeon and fowl the legs and the two outer toes are heavily feathered,
so that, in the trumpeter pigeon, they appear like little wings. In the
feather-legged bantam, the 'boots,' or feathers, which grow from the
outside of the leg, and generally from the two outer toes, have, according
to the excellent authority of Mr. Hewitt, been seen to exceed the
wing-feathers in length, and in one case were actually nine and a half
inches in length! As Mr. Blyth has remarked to me, these leg-feathers
resemble the primary wing-feathers, and are totally unlike the fine down
which naturally grows on the legs of some birds, such as grouse and owls.
Hence it may be suspected that excess of food has first given redundancy to
the plumage, and then that the law of homologous variation has led to the
development of feathers on the legs, in a position corresponding with those
on the wing, namely, on the outside of the tarsi and toes. I am
strengthened in this belief by the following curious case of correlation,
which for a long time seemed to me utterly inexplicable,--namely, that in
pigeons of any breed, if the legs are feathered, the two outer toes are
partially connected by skin. These two outer toes correspond with our third
and fourth toes. Now, in the wing of the pigeon, or any other bird, the
first and fifth digits are wholly aborted; the second is rudimentary, and
carries the so-called 'bastard wing;' whilst the third and fourth {182}
digits are completely united and enclosed by skin, together forming the
extremity of the wing. So that in feather-footed pigeons not only does the
exterior surface support a row of long feathers like wing-feathers, but the
very same digits which in the wing are completely united by skin become
partially united by skin in the feet; and thus, by the law of the
correlated variation of homologous parts, we can understand the curious
connexion of feathered legs and membrane between the outer toes."
Irregularities in the circulating system are far from uncommon, and
sometimes illustrate this homological tendency. My friend and colleague Mr.
George G. Gascoyen, assistant surgeon at St. Mary's Hospital, has supplied
me with two instances of symmetrical affections which have come under his
observation.
Public-domain text, read in full here on John Shaqi.
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