On the Variation of Species, with Especial Reference to the Insecta: Followed by an Inquiry into the Nature of GeneraWollaston, Thomas Vernon
Science
On the Variation of Species, with Especial Reference to the Insecta: Followed by an Inquiry into the Nature of Genera
Wollaston, Thomas Vernon
Insects; Variation (Biology)
As regards positive structure, indeed, we can have but few
observations to communicate,--seeing that the limbs and appendages
themselves are usually of so constant a nature, that disturbing
agencies have little or no power to divert them from their typical
states. Still, there are occasional facts on record, which would tend
to prove that even these are not altogether exempt from the deranging
force of certain contingencies from without: the number of the
antennal joints, for instance, in the tribes where those organs are
multiarticulate, is said to vary; but how far this may be dependent on
physical influences, I am not in a position to decide. The connateness
of the elytra, again, is a character which we may at any rate define
as _sub_-structural; and this I have myself noticed, at times, to
fluctuate, according to the circumstances and conditions of the
respective localities in which the particular species obtain. Such is
eminently the case with the universal _Harpalus_ (the _H. vividus_,
Dej.) of the Madeiran Group. Speaking of this peculiarity, in my
volume on the Coleoptera of those islands, I made the following
remarks: "But perhaps its most singular character, and in which it
differs from every other _Harpalus_ with which I am acquainted,
consists in the tendency of its elytra to become united or soldered
together. I say 'the tendency,' because it is not always the case that
they are joined (which, since the law exists at all, is perhaps the
more remarkable), although in most instances, especially in localities
much exposed and but slightly elevated above the sea-shore, they are.
I have examples, however, from the upper as well as the lower regions,
in which both states are represented; and others again in which the
elytra are only partially connected, being free at the apex though
firmly attached towards the scutellum. In every instance, however,
even where they are united throughout their entire length, a little
force will succeed in separating them, showing their structure, as I
have indicated in the diagnosis, to be _sub_-connate rather than
connate. But that it does require force to effect the disjunction,
when they are really in the condition described, is proved to a
demonstration to any one who has seen the _remains_ of the insect
beneath the slabs of stone on many of the small adjacent islands where
it most abounds, or drifting about over the surface of the
rocks,--under which circumstances I have observed them in immense
numbers, apparently the accumulation of two or three generations,
which the violence of the elements had not been able to sever. It is
rare in the sylvan districts to find them joined; nevertheless such is
sometimes the case,--thus proving that the peculiarity is not actually
essential, but merely one which it is the tendency of the species to
assume, and which is more developed in some specimens, and under
certain conditions, than in others.[46]"
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