On the Variation of Species, with Especial Reference to the Insecta: Followed by an Inquiry into the Nature of GeneraWollaston, Thomas Vernon
Science
On the Variation of Species, with Especial Reference to the Insecta: Followed by an Inquiry into the Nature of Genera
Wollaston, Thomas Vernon
Insects; Variation (Biology)
In like manner, the genus _Tarphius_ is distinctively Madeiran. I have
detected nearly twenty well-defined species of it in that group; yet,
out of so large a number, two only have occurred beyond the central
island. Now the _Tarphii_ are, also, wingless; and creatures of very
sluggish propensities,--scarcely ever stirring from the masses of
loose rotting timber which they so assimilate in hue, and to the under
sides of which they affix themselves, day and night. Although
difficult to investigate in their precise economy, it is extremely
probable (may I not say, certain?) that some important and peculiar
office is assigned to them in the remote upland districts to which
they exclusively belong: and there cannot be any question, to a person
who has studied them carefully on the spot, but that the region which
they now inhabit is the actual area of their primaeval appearance on
this earth. Many kindred species may of course have been lost, during
those gigantic subsidences which caused the Madeiras to be shaped out,
and to tell their tale above the waves as ruins of an ancient land;
yet our existing cluster of forms could not have wandered far at that
early period, from the Serras and ridges of their birth,--perhaps not
_so_ far indeed (considering the limited bounds within which they are
now confined, and that time should in reality have increased their
range rather than diminished it) as they have succeeded in doing at
the present day. Hence we may reasonably conclude, that Madeira proper
is an example of what we have alluded to in a preceding page,--namely,
of the accidental retention, during a vast downward movement, of a
nucleus of small specific areas of colonization, the colonizers of
which _had not extended elsewhere_. But I stated, that two of the
above-mentioned _Tarphii_ have occurred beyond the central mass. It is
in Porto Santo that they make their appearance; nevertheless, since
one of them is apparently peculiar to that island, it is only the _T.
Lowei_, Woll. (an insect of a different, and more active, nature than
the rest) which has violated that _local exclusiveness_ which would
seem to be almost a generic character, as it were, of its allies. That
species, however, both in its manners and aspect, recedes materially
from the remainder. Although, like them, nocturnal in its habits, it
is able to run with considerable velocity; and, instead of attaching
itself to the blocks of putrefying wood, which both fall and decay _in
situ_ on those elevated tracts, it hides within the bunches of
_Evernia scopulorum_ and _prunastri_ which clothe the trunks of living
trees, and fill up the crevices of the weather-beaten peaks. Hence,
when contrasted with its comrades, we can easily understand how the
varied processes of accidental transportation would operate to
increase the range of a creature which differs so essentially, in many
respects, from them. It is indeed, not unfrequently, brought down, at
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