On the Variation of Species, with Especial Reference to the Insecta: Followed by an Inquiry into the Nature of GeneraWollaston, Thomas Vernon
Science
On the Variation of Species, with Especial Reference to the Insecta: Followed by an Inquiry into the Nature of Genera
Wollaston, Thomas Vernon
Insects; Variation (Biology)
_Deucalion_, also, may be quoted in
support of this twofold hypothesis, of the direction, and the
slowness, of the former migratory movements. It is an apterous genus,
and of eminently sluggish habits; and what is the consequence?--we
have a very remarkable species (the _D. oceanicum_, Woll.) on one of
the rocks of the Salvages, whilst another (the _D. Desertarum_, Woll.)
has been isolated on the two southernmost islands of the Madeiran
Group; and of so sedentary a nature is this last, that, although
physically unimpeded, it has not, even to this day, overrun the
diminutive areas on which, when the surrounding region was submerged,
it was originally saved from destruction. So strongly indeed was this
fact impressed upon me, when I first detected it, that I shall perhaps
be excused for recapitulating _in extenso_ the few reflexions which
then suggested themselves to my mind. "There is no genus, perhaps,
throughout all the Madeiran Coleoptera, more truly indigenous than
_Deucalion_. Confined apparently, so far as these islands are
concerned, to the remote and almost inaccessible ridges of the two
southern Dezertas, it would seem to bid defiance to the most
enthusiastic adventurer who would scale those dangerous heights. Its
excessive rarity, moreover, even when the localities are attained,
must ever impart to it a peculiar value in the eyes of a naturalist;
whilst its anomalous structure and sedentary[60] mode of life give it
an additional interest in connexion with that ancient continent, of
which these ocean ruins, on which for so many ages it has been cut
off, are the undoubted witnesses. Approximating in affinity to
_Parmena_ and _Dorcadion_, yet presenting a modification essentially
its own, it becomes doubly important in a geographical point of view;
and it was therefore with the greater pleasure that I lately
received a second representative, from the distant rocks of the
Salvages,--midway between Madeira and the Canaries. Differing widely
in specific minutiae, yet agreeing to an identity in everything
generic, they offer conjointly the strongest presumptive evidence to
the _quondam_ existence of many subsidiary links (long since lost, and
radiating in all probability from some intermediate type) during the
period when the whole of these islands were portions, and perhaps very
elevated ones, of a vast continuous land. * * * * * The _Deucalion
Desertarum_ is of the utmost rarity, the only two[61] specimens which
I have seen having been captured (the first by myself, in 1849; and
the second by the Rev. R. T. Lowe, in 1850) on the respective summits
of the Middle and Southern Dezertas. So local indeed does it seem to
be, that it, apparently, has not extended itself even over the Dezerta
Grande (where there are no external obstacles to bar its progress);
but retains the very position which in all probability constituted its
original centre of dissemination at the remote period of time when
this ancient continent received its allotted forms.
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