On the Variation of Species, with Especial Reference to the Insecta: Followed by an Inquiry into the Nature of GeneraWollaston, Thomas Vernon
Science
On the Variation of Species, with Especial Reference to the Insecta: Followed by an Inquiry into the Nature of Genera
Wollaston, Thomas Vernon
Insects; Variation (Biology)
As regards Professor Forbes's views of the creation of a vast
continent (reaching far into the Atlantic[66]) at the close of the
miocene epoch, through the upheaved bed of a shallow miocene sea,--a
region moreover of such an extent as to have connected the various
island groups between the Fucus bank and the shores of the Old World,
not only with each other, but with a Mediterranean province, Asturias,
and even the south-west of Ireland,--I must be content to pass them
by, hazarding only a few crude and desultory remarks. So large a
question, indeed, cannot be safely handled without a corresponding
amount of data, in all departments of natural science, to reason
from,--which I do not possess: still, if a speculation from
entomological premises, _per se_, be not altogether worthless, I would
point to the conclusions (lately adverted to) which my Madeiran
researches have forced upon me, concerning the _direction_ of the
former insect migrations,--inferences which are, from first to last,
of necessity erroneous, if the requisite medium for transit (into
South-European latitudes, at all events) be a mere conjecture or
romance. Such a notion, however, I would not for a moment
entertain,--for there is too much direct evidence in support of
distinct epochs of diffusion, to allow of any hypothesis, when
endeavouring to account for the phaenomena which we now behold, to
supersede the assumption of a once continuous tract. No matter if we
be compelled to suppose, whilst attempting to interpret what we see,
that the disseminating current has flowed in exactly opposite courses,
at different and remote periods, over the surface of that ancient
land,--seeing that the _fact_ (if such in reality it be) remains
untouched, that _the land itself is_ at any rate _there_. I am not,
however, prepared to assert that the opinion at which I had
independently arrived, from the insect statistics, does positively
require a northerly prolongation of that area beyond the line of the
central Mediterranean districts; yet, after making every possible
allowance for accidental introductions since the subsidences have
taken place, there is still left a large residuum which I am convinced
can never be explained (unless the doctrine of specific centres be a
myth) except through the means of ordinary and regular migration over
an unbroken continent. Nevertheless, though I would not presume, from
insufficient material, to insist upon an extension of this Atlantic
region into higher latitudes than those which I have just referred to,
I must express my individual belief that, the more the subject is
examined, with reference to the distribution of the Annulosa, the
less will Professor Forbes's idea suffer from the inquiry. In the
'Insecta Maderensia,' I have already thrown out a few scattered hints
which bear on this immediate consideration; and, since no subsequent
reason has induced me either to withdraw or modify them (but rather
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