On the Variation of Species, with Especial Reference to the Insecta: Followed by an Inquiry into the Nature of GeneraWollaston, Thomas Vernon
Science
On the Variation of Species, with Especial Reference to the Insecta: Followed by an Inquiry into the Nature of Genera
Wollaston, Thomas Vernon
Insects; Variation (Biology)
Conceding, therefore, this distinction between barriers of a primaeval
and more recent character, it is not difficult to understand why the
opposite sides of an alpine chain, as well as countries separated by
the sea, should display different phaenomena from each other. On the
contrary indeed, if we could feel satisfied that no means of
accidental transportation had operated to take them there, and that
the animals themselves were incapable of enduring great diversities of
temperature, and other contingencies; we should be startled to
discover creatures specifically identical in such regions,--so long at
least as the doctrine of unique centres of radiation formed part of
our zoological creed. We must not, however, be too hasty in
questioning (if I may be pardoned for the completion of a metaphor of
which I thoroughly disapprove) this article of our faith, through the
occurrence of similar beings in areas between which there exist
barriers, both primary and well-defined; for the methods of diffusion
are so complicated and numerous, that, even where human agency (that
most important of elements) is not concerned, what at first sight may
frequently appear to be impossible becomes clear enough when more
critically inquired into. Some species, we know, are gifted with
greater powers for horizontal and vertical progression than their
comrades, and can (though they are doubtless exceptions to the general
rule) pass through extremes of atmosphere sufficient to render even
lofty mountain summits no obstacles to _them_. Others, as the
_Calosoma Syncophanta_ of Europe, have been stated to traverse the
ocean unhurt[70]; and I believe that many do at times accidentally
arrive, in a half-drowned state, especially after boisterous weather,
across channels of considerable breadth. Mr. Kirby, on examining the
marine _rejectamenta_, during one of these apparent occurrences, along
the Suffolk coast, writes as follows: "Whether the insects I observed
upon the beach, wetted by the waves, had flown from our own shores,
and, falling into the water, had been brought back by the tide; or
whether they had succeeded in the attempt to pass from the continent
to us, by flying as far as they could, and then falling had been
brought by the waves, cannot certainly be ascertained; but Kalm's
observation inclines me to the latter opinion[71]." And Sir Charles
Lyell remarks:--"Exotic beetles are sometimes thrown on our shore,
which revive after being drenched in salt water[72]." Nor should we
forget that chance agencies of every description, which we are too apt
to overlook, are daily at work (and have been so since, at any rate,
the last creative epoch) to transport these variously organized beings
beyond their original spheres. Sometimes they are carried on, or
within, the bodies of larger animals, which is especially the case
with the parasitic tribes; at others on floating trunks of trees, and
casual substances of divers kinds, which are able to resist for a
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