Physiology: The Science of the BodyMartin, Ernest G.
Science
Physiology: The Science of the Body
Martin, Ernest G.
Physiology
All that has been said about the nervous system thus far has applied to
its duty of controlling the activities of skeletal muscles; namely, the
muscles by which the body carries on its principal motions. These, as
already stated in Chapter VII, are not the only kind of muscles in the
body. There are also the smooth muscles and heart muscle. There is also
another kind of functional metabolism besides the metabolism of muscular
motion over which the nerves have control. This is the metabolism of
secretion, or the manufacture of special chemical substances by gland
cells. In later sections of the book we shall hear more about what the
glands do, but here we have to say something about the way they are
controlled through the nervous system. We speak of them along with the
smooth muscles, because it happens that the same nervous arrangements
are found in both cases; the smooth muscles and the glands have a
precisely similar kind of nervous control. In appearance the nerves
which lead from the central nervous system to the smooth muscles and the
glands do not differ much from other nerves. They are composed of nerve
cells, just as are all the other nerve pathways. It happens to be true
that the sheaths which inclose the axons of these particular nerves
usually have little or no white material in them, so that nerve trunks
which are made up of these fibers look gray instead of white, as do the
other nerves. Just why there should be this difference, we do not know.
Of more importance than this difference in appearance is the fact that
the connection from the central nervous system to the smooth muscle cell
and the gland cell in no case is made up of a single nerve cell, but
always of a chain of two. In the case of nerve paths to skeletal muscles
the cell body of the motor nerve cell lies either in the spinal cord or
the brain and its axon extends all the way from there directly to the
muscle fibers. In the case of smooth muscles and glands a connecting
nerve cell extends out from the central nervous system some distance and
makes its nerve junction with the motor nerve cell part way between the
central nervous system and the muscle or gland. These connecting cells
are much branched, as are all connecting cells; also the motor cells
which connect with smooth muscles or glands are very much branched; the
result is that a single nerve cell starting from the spinal cord finally
connects with a large number of smooth muscle cells or gland cells. We
do not have, then, in these organs the fine subdivision of control that
there is in the skeletal muscles, for a single skeletal motor cell in
the spinal cord connects with one or, at the most, a very few muscle
fibers. If we consider the uses to which smooth muscles are put, we
shall see that finely subdivided control is not important, as it is in
the case of the skeletal muscles, where the movements have to be graded
very accurately. Smooth muscles are present in the stomach and
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