After the physicist has arrived at the idea that the semi-circular canals are the organ of sensation of rotation or of angular acceleration, he is next constrained to ask for the organs that mediate the sensation of acceleration noticed in forward movements. In searching for an organ for this function, he of course is not apt to select one that stands in no anatomical and spatial relation with the semi-circular canals. And in addition there are physiological considerations to be weighed. The preconceived opinion once having been abandoned that the _entire_ labyrinth is auditory in its function, there remains after the cochlea is reserved for sensations of tone and the semi-circular canals for the sensation of angular acceleration, the vestibule for the discharge of additional functions. The vestibule, particularly the part of it known as the sacculus, appeared to me, by reason of the so-called otoliths which it contains, eminently adapted for being the organ of sensation of forward acceleration or of the position of the head. In this conjecture I again closely coincided with Breuer. That a sensation of position, of direction and amount of mass-acceleration exists, our experience in elevators as well as of movement in curved paths is sufficient proof. I have also attempted to produce and destroy suddenly great velocities of forward movement by means of various contrivances of which I shall mention only one here. If, while enclosed in the paper box of my large whirling apparatus at some distance from the axis, my body is in uniform rotation which I no longer feel, and I then loosen the connexions of the frame _rr_ with _R_ thus making the former moveable and I then suddenly stop the larger frame, my forward motion is abruptly impeded while the frame _rr_ continues to rotate. I imagine now that I am speeding on in a straight line in a direction opposite to that of the checked motion. Unfortunately, for many reasons it cannot be proved convincingly that the organ in question has its seat in the head. According to the opinion of Delage, the labyrinth has nothing to do with this particular sensation of movement. Breuer, on the other hand, is of the opinion that the organ of forward movement in man is stunted and the persistence of the sensation in question is too brief to permit our instituting experiments as obvious as in the case of rotation. In fact, Crum Brown once observed while in an irritated condition peculiar vertical phenomena in his own person, which were all satisfactorily explained by an abnormally long persistence of the sensation of rotation, and I myself in an analogous case on the stopping of a railway train felt the apparent backward motion in striking intensity and for an unusual length of time.