The local species of _Agelaius_ show similarly capricious distinctions.
_A. phoeniceus_ is a widely spread species, found over a great part of
North America. The male is black with red-orange bars on the wings, but
the female is somewhat thrush-like in colour. In the island of Porto
Rico there is a form called _xanthomus_, in which _both sexes_ are like
the males of the mainland. A similar species called _humeralis_, also
with both sexes male-like, lives in Cuba. The island of Cuba, curiously
enough, has also a distinct species named _assimilis_, in which the
female is a dull black all over, though the male is like the mainland
type.
So also may local races differ in respect of variability. _Argynnis
paphia_, the Silver Washed Fritillary, through a great part of its
distribution has only one female form. In the English New Forest a
second female form, _valesina_, co-exists with the ordinary _paphia_
female. But in the southern valleys of the Alps the _valesina_ female
is much the commoner of the two, and indeed in some localities where
the species is abundant, I have seen no _paphia_ females in many days
collecting.
The beetle _Gonioctena variabilis_ furnishes an illustration of a
comparable phenomenon affecting the male sex. In 1894 and 1895 I
studied the curious colour variations of this species especially in the
neighbourhood of Granada, and Mr. Doncaster ten years later repeated the
observations on the same ground, and also collected the insect in other
places in the south of Spain. The distinctions are not easy to give in
words and the reader is referred to the colour plate accompanying my
paper.[5] The essential fact is that the males commonly have the elytra
_red with black spots_ and the females for the most part have greenish
grey elytra with black stripes. In some localities a large minority of
males closely resemble the female type, being identical in colour and
then only distinguishable by structural differences. In two Granada
localities I found the proportion of such males quite different. In the
Darro valley about 38 per cent. (in 718) were of this feminine type,
but on the hills some 300 feet above only 19 per cent. (in 3,230) were
like the females. At Castillejo, not far from Toledo I found no such
male in 75 specimens.
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