Species which are uniform in some localities may be polymorphic in
others. Such a phenomenon is well exemplified by the orchid _Aceras
hircina_. Of this species distinct varieties had previously been known
in Germany, but Gallé[9] has lately given a detailed account of a
number of most diverse forms found growing in a district of Eastern
France. Without reference to his plates it is impossible to give any
adequate conception of the profusion of types which the flowers of
the species there assume. In some the lip is elongated to many times
its usual length, twisting and dividing in a fashion suggesting some
of the strangest of the Tropical Orchids. In others the labellum
and the lateral petals are all comparatively short and wide (Fig.
13). Intermediates, combining these qualities in various degrees,
were abundant, and the condition of the species, which was the only
representative of the genus in the locality, recalls the extreme
polymorphism of many of the Noctuid Moths.
[Illustration: FIG. 13. Various forms of _Aceras hircina_. (After
Gallé.) This figure only shows a few of the more striking forms
illustrated in Gallé's plates.]
Somewhat comparable variability has been seen in another Orchid genus
_Ophrys_. In Great Britain the species _apifera_, _aranifera_ and
_muscifera_ though variable are fairly distinct, but Moggridge has
published two series of plates[10] showing a very different state of
things as regards the _Ophrys_ population of the Riviera. Here the
outward diversity is such that the ordinary specific names cannot be
applied with any confidence and the limits of the species are quite
uncertain. It may well be supposed that these Riviera plants are
interbreeding, and indeed we may safely assume that they are. It is,
however, to be remembered that Darwin showed _apifera_ in this country
to be habitually self-fertilised, so that the different behaviour on
the Riviera may itself constitute a local peculiarity. Moreover it is
to be gathered from Moggridge's account that in the districts which he
examined the condition was not to be described by the statement that
our three types were there co-existing and hybridising, but rather we
should say that the population was polymorphic, containing these three
types amongst others. Conchologists are aware that on the Dogger Bank
_Modiola_ attains a size unparalleled elsewhere. The same is true of
the sponges _Grantia compressa_ and _Grantia ciliata_ in the estuary
of the Orwell.[11] Conversely, as we know so well in the case of Man,
dwarf races occur in several special localities. Such examples may be
multiplied indefinitely.
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