Taking the evidence as a whole, we may say that it fairly suggests the
existence of some connection between modern urban developments and the
appearance and rise of the melanic varieties. More than that we cannot
yet affirm. It is a subject in which problems open up on every side,
and all of them are profitable subjects for investigation. Unhappily
such animals are difficult to rear successfully in captivity for many
generations, owing to their extreme liability to disease. Not the least
interesting feature of the melanics is the fact that the black varieties
provide about the best and clearest example of a new dominant factor
attaching itself to a wild species in recent times. None of the cases
are satisfactorily recorded or analysed as yet, but the evidence is
clear that _doubledayaria_ is a dominant to its type, and in several
other dark varieties, though the pigment deposited is not black, the
records show that the increased amount of the pigment almost certainly
is due to a positive factor. Of this, _Hemerophila abruptaria_ is a
good example.[24] There are some irregularities in the results, but
taken together they leave little doubt that the dark brown variety is a
dominant and the light, yellowish brown a recessive.
A curious parallel to the rise of the melanic moths in England
is provided by the case of the Honey-creepers or Sugar-birds, in
certain West Indian islands.[25] These birds of the genus _Coereba_
(_Certhiola_) range from Southern Mexico to the Northern parts of
South America and through the whole chain of the West Indian islands
and Bahamas except Cuba. There are numerous local forms, and many of
the islands have types peculiar to themselves, as is usual in such
cases. Some of the types or species range through several islands, but
according to Austin Clark[26] no island has more than one of them.
Cory[27] reckoned twelve such species within the Antillean region. They
are small birds about the size of a nuthatch with a general colouring of
black, yellow, and white. From the island of St. Vincent the Smithsonian
Institution received in the late seventies of last century several
completely black specimens in addition to two of the usual type of
colouring. The black were described by W. N. Lawrence as _atrata_, and
those marked with the usual yellow and white were called _saccharina_.
The collector (Mr. F. A. Ober) reported that the black form was common,
and that the _saccharina_ form was rarer. Lawrence remarks, "Had there
been only a single example (of the black form) I should have considered
it as probably a case of abnormal colouring, but it seems to be a
representative form of the genus in this island."[28] There is of course
no doubt of the correctness of the view taken by Austin Clark that
"_atrata_" is a black variety. The black bird is in every respect, other
than colour, identical with _saccharina_, and it is even possible to
detect a greenish colour in the areas which would normally be yellow,
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