It is generally supposed that phenomena like those now so well
established in the case of _Colaptes_ are very exceptional, and as has
already been stated a number of circumstances must combine in order
that they may be produced. I suspect however that the examples are
more numerous than is commonly thought. In all likelihood the three
forms _Sphyropicus varius_, _nuchalis_ and _ruber_ are in a very
similar condition though the details have not, so far as I know, been
worked out. A complex example which is closely parallel to the case of
_Colaptes_ was described by F. M. Chapman[9] at the same date as Allen's
work. This is the case of _Quiscalus_, the Grackles, which in the North
American Continent have three fairly distinct forms which Chapman speaks
of as _Q. aeneus_, _Q. quiscula_, and _Q. quiscula aglaeus_. The birds
are all, so far as pigment is concerned, dark blackish brown, but the
head and mantle have superposed a metallic sheen of interference-colours
which in the various forms take different tints, bluish green, bronze
green, or bronze purple. The details are complicated and difficult
to appreciate without actual specimens, but the two common types are
sufficiently distinct. The birds inhabit the whole area east of the
Rockies, _quiscula aglaeus_ occupying Florida and the Southern States
southwest of a band of country about a hundred miles broad extending
roughly from Connecticut to the mouth of the Mississippi; and _aeneus_
taking the area north and west of this band. In discussing this case
Chapman expresses the same view as Allen does in the _Colaptes_ case,
that there are two distinct populations, substantially fixed, and
that the band of country in which they meet each other has a mongrel
population, with no consistent type, but showing miscellaneous
combinations of the character of the two chief types.
The warblers of the genus _Helminthophila_ provide another illustration
which has points of special interest. The two chief species are _H.
pinus_, which has a yellow mantle and lower parts, white bars on the
wings, a black patch behind the eyes and a broad black mark on the
throat; and _H. chrysoptera_ with dark grey mantle and pale whitish grey
lower parts, yellow bars on the wings, and grey marks on cheeks and
throat where _pinus_ has black. These two birds are exceeding distinct,
and in addition their songs are quite unlike. _H. pinus_ ranges through
the eastern United States up to Connecticut and Iowa. _H. chrysoptera_
is a northern form extending down to Connecticut and New Jersey. Both
are migrants.
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