FIG. 4. "Brewster's Warbler," male; another of the integrading forms.
FIG. 5. _Helminthophila chrysoptera_, male.
FIG. 6. _Helminthophila chrysoptera_, female.]
Allen[11] has described another excellent American example, the Tits of
the group _Baeolophus bicolor-atricristatus_. The form _bicolor_ belongs
to the eastern States and ranges from the Atlantic coast to the Great
Plains, and _atricristatus_, of east Mexico, extends from Vera Cruz to
central Texas. In southern and central Texas the breeding ranges adjoin,
and in this country various intermediates occur. The chief types differ
in two main points.
_B. bicolor_
Forehead varies from deep _black_
to dull black, suffused with
rusty brown.
Crown and crest _grey_,
slightly darker than the back.
_B. atricristatus_
Forehead _white_ to buffish white.
Crown and crest _black_, abruptly
contrasting with the back.
The intergrades between the two have, as usual, received specific names.
A detailed description is given by Allen, from which it appears that
the gradation is very complete. In one case a series of 16 adults were
all intermediates. It is not stated whether the collector took these at
random, but from the local lists it is clear that the types are found
not far away from the place where the intergrades were shot.
Another very striking case is that of the Tanagers, of the genus
_Rhamphocoelus_. In this group there are several local forms which
are related to each other in remarkable ways. The forms known as
_passerinii_ and _icteronotus_ exhibit the clearest phenomena of
intergradation. The species _passerinii_ has a brilliant scarlet
and black male, and it inhabits Honduras and Nicaragua. Proceeding
southwards along the isthmus we find next _costaricensis_ which has
a male like that of _passerinii_ (but a female with more orange than
the olive-grey female of _passerinii_). Next we come to Panama which
is occupied by _icteronotus_, sharply distinguished from _passerinii_
by the fact that the _scarlet is replaced by lemon-yellow_. This same
_icteronotus_ occurs again as a pure type in Ecuador and many other
parts of South America; but Colombia, _between Panama and Ecuador_,
contains scarlets like _passerinii_, yellows like _icteronotus_, and
various intergrades of several shades of orange. The _passerinii_ males
from Nicaragua are indistinguishable from those of Colombia, and the
_icteronotus_ of Ecuador are the same as those in Panama. The orange
intergrades, doubtless heterozygous forms, though collected at the
same locality (Medellin in Colombia) as several pure yellows and pure
scarlets, are in the British Museum series sorted out as a separate
species under the name _chrysonotus_! Complications are introduced by
the relations of these forms to another named type, _flammigerus_, but
we may for our purpose leave that out of consideration, and say that the
order of geographical sequence from Honduras to Ecuador is (1) scarlet,
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