From this particular discussion I omit reference to those examples in
which the permanently established types are obviously associated with
special conditions of life. Where considerable climatic differences
exist between localities, or when we pass from South to North, or from
the plains into Alpine levels we often find that in correspondence
with the change of climate there is a change in the characteristics of
a species common to both. When I say "species" in such a connection I
am obviously using the term in the inclusive sense. Some would prefer
to say that in the two sets of conditions two _representative species_
exist. Whichever expression be preferred it is plain that such examples
present another phase of the problem we have been just considering, and
in them also we have an opportunity of observing the consequences of
the overlap of two closely related types, but there are advantages in
considering them separately. In the examples hitherto given, with the
possible exception of the Papilios,[15] the two fixed types severally
range over so extensive a region that it may fairly be supposed that
in the different parts they are subject to considerable diversities of
climate. There is no outstanding difference that we know distinguishing
the habitats of the two forms; but in comparing Alpine with Lowland
forms, or essentially northern with essentially southern forms we do
know an external circumstance, temperature, that may reasonably be
supposed to have an influence, direct or indirect, on the population.
FOOTNOTES:
[1] J. A. Allen, _The North American Species of the Genus Colaptes,
Considered with Special Reference to the Relationships of C. auratus and
C. cafer_. Bull. Am. Mus. Nat. Hist., IV, 1892.
[2] For a case in which a red-headed female × a black-headed male gave a
black-headed female and a red-headed male, see _Avian Mag._, N. S., IV,
pp. 49 and 329
[3] The other variations of this bird are also interesting and
important. The normal male has a red head and a red throat. The female
has a red head and a white throat, but varieties of the female are known
with a black head, thus again illustrating the change from black to
red. It should be noted that this is not a mere retention of a juvenile
character, but, as the birds mature, the red feathers come up, or as an
exception, the black. There is also a western species, _ruber_, in which
both sexes have a great extension of red, and are alike. The male of
_nuchalis_ intergrades with this type, but the female does not.
[4] Dr. W. Brewster, for example, has a remarkable specimen of the Teal
(_Nettion carolinense_) with a white collar strongly developed at the
front and sides of the neck, in a place where the normal has no such
mark.
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