Another form in which I tried to investigate the same problem is
_Coenonympha arcania_, which has one Alpine form known as _darwiniana_,
and another, _satyrion_. In calling _satyrion_ a form of _arcania_ I
follow Staudinger and other authorities, but I have never been quite
satisfied that it should be so regarded. The differences between
_arcania_ and _darwiniana_ are essentially differences of degree; _C.
arcania_ occurs in places where there is cover, and reaches up the
valleys usually as high as the mixed woods of deciduous trees, which
is about 2,500 feet. The variety _darwiniana_, on the contrary, is an
insect of treeless hillsides, and I regard it as a dwarf and possibly
a stunted form. It would not greatly surprise me to find that with
the application of good conditions _arcania_ could be raised from
_darwiniana_ eggs, or that if _arcania_ larvae were starved they might
give rise to _darwiniana_ butterflies. I have been unsuccessful in
trying to rear the species, having lost the larvae by disease. Usually
one does not catch _arcania_ and _darwiniana_ on the same ground, and
as _Festuca ovina_--a typically hill-side grass--is a common food-plant
of _darwiniana_ there can be little doubt that _arcania_ feeds on
some other grass, probably woodland species. Colonies of _arcania_ of
varying size and brightness are commonly found, and though a sample of
_arcania_, finely grown, from a warm Italian wood, presents a striking
contrast with _darwiniana_ from an Alpine pasture, one certainly may get
samples which fill all the gradations. Generally the sample from a given
locality is fairly homogeneous.
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