Schröder's second example is not more convincing, in my opinion, though
Semon regards it as one of the most important pieces of evidence. It
concerns a leaf-rolling moth, _Gracilaria stigmatella_, the larva of
which is said normally to make its house by bending over the _tips_ of
the sallow leaves on which it feeds. Schröder placed larvae on leaves
from which the tips had been cut, and these larvae made their houses by
rolling over the _sides_ of the leaves. Their offspring were again fed
on leaves without tips, and as before, they rolled in the leaf-margins
either on one side or both. The offspring of this second generation were
then fed on entire leaves. There were 19 houses made by these (?19)
larvae, and of them 15 were normal, made by folding down the tips of
the leaves, while 4 were abnormal, made by rolling in the leaf-margins.
Schröder says that in nature he has only twice seen abnormal houses; but
it is clearly essential not only that the frequency of such variability
in nature should be thoroughly examined, but also that we should know
whether when the species is bred in captivity these irregularities of
behaviour do or do not occur when the larvae are fed on uninjured leaves.
The famous case of Schübeler's wheat is revived by Semon. The story will
be familiar to most readers of the literature of the subject. Briefly
it is that annuals, especially wheat and maize, raised from seed in
Central Europe take more time in coming to maturity and ripening than
similar plants raised in Norway, where the summer days are much longer.
The received account is that he imported seed especially of maize and
of wheat from Central Europe to Norway and found that in successive
years the period of growth and ripening was increasingly reduced. After
two generations seed of the accelerated wheat was sent back to Breslau
where it was grown, and was found to ripen rather more slowly than in
Norway, but much more quickly than the original stock had done. The
facts recorded by Schübeler[6] are that he received seed from Eldena,
which is on the Baltic near Greifswald. The variety is described as
"_100 tägiger Sommer Weizen_," but no more exact record of its behaviour
in Germany is given. This wheat, grown at Christiania in 1857, took
103 days to harvest. Its seed was again grown in Christiania in 1858,
and took 93 days, and sown again in 1859 it took only 75 days, 28 days
less than in the first year of cultivation in Norway. Seed of the 1858
crop was sent to Breslau, and grown there by Roedelius in 1859; it
took 80 days. Evidently before such a record can be used as proving an
inheritance of acquired characters numbers of particulars should be
forthcoming. The view that Johannsen has taken is that the result was
probably due to unconscious selection of the earlier individuals among a
population consisting of many types of various compositions. Some effect
may no doubt be ascribed to that cause, but I cannot think that alone
it would account for the results.
Public-domain text, read in full here on John Shaqi.
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