The reciprocal cross was made between an _abnormal female_ and a _normal
male_. In this case the offspring were abnormal in their behaviour; but
the second generation bred from them showed three quarters abnormal and
one quarter normal.
Certain details as to numbers and sexes of the various families bred
in the course of this amazing experiment are given in a subsequent
publication.[15] This later paper goes somewhat fully into the question
of the difference in behaviour between the normal and modified
individuals, describing the ways in which the males and females
possessing the acquired character could be recognised from the males and
females which were normal, but in this account I find no reference to
the development of the "_Brunftschwielen_"--the horny pads on the hands
of the males. As these structures would be of special value in such a
diagnosis the omission of any allusion to them calls for explanation.
Kammerer claims the evidence as proof of Mendelian segregation in
regard to an acquired character, the first example recorded. Pending a
repetition of the experiments there is no more to be said.
2. _The Mode of Reproduction of Salamandra atra and
maculosa._[16]--_Salamandra maculosa_, the common lowland form, with
yellow bands or spots, deposits its young in water, generally as
gill-bearing tadpoles, with a wide, swimming tail, though occasionally
they are born still enclosed in the egg-capsule out of which they soon
hatch. Spawning extends over a considerable period, often many weeks,
and during the season one female may bear more than 50 young.
_S. atra_, the black Alpine form, produces its young on land. They are
born without gills, ready to breathe air, and with the rounded tail
of the adult. These differences may, as Kammerer says, naturally be
regarded as adaptations to the Alpine conditions. Moreover, the female
bears _only two_ young in a season, and this reduction in the number
must be taken to be a consequence or condition of viviparity. There are
many eggs in the ovary, but all except the two which are destined to
develop degenerate and form a yolk-material on which these two survivors
feed.
Kammerer gives a long account of the various conditions to which he
subjected both species. The treatment was complicated in many ways, but
the essential statements are, as regards _S. maculosa_, that when no
water was provided in which the young might be born, they were dropped
on land, larger and in a later stage of development and of a darker
colour than is normal; that the larvae so born gradually diminished
in number until only two were deposited in each breeding-period;
that dissection showed that the other ova degenerated to form a
yolk-material. The larvae so produced reached maturity. The summary of
results describes their behaviour, stating that they produced:
Public-domain text, read in full here on John Shaqi.
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