In continuation[25] of the experiments on the colour of _S. maculosa_
Kammerer publishes an account of elaborate experiments in grafting
ovaries of the various forms, modified and unmodified, into each other,
and describes the offspring which followed. Before pursuing this part of
the inquiry I am disposed to wait until the earlier steps have been made
much more secure than they yet are.
More recently Kammerer has published similar statements in regard to the
inheritance of characters induced in various lizards by keeping them in
abnormal temperatures, high and low. The changes induced affected in
some species the colours, in others the reproductive habits. Respecting
these examples I feel the same scepticism that I have indicated in
regard to the others, somewhat heightened by the fact that insufficient
evidence is given both regarding the behaviour of these various species
in captivity when not subjected to abnormal temperatures, and in the
wild state.
Respecting this part of the evidence Mr. G. A. Boulenger has lately
published a criticism[26] from which I extract the following passages.
Referring to a previous note[27] on the question of the melanism of the
various insular forms of _Lacerta muralis_ he writes: "I also alluded
(_l. c._) to the theories that have been propounded to explain the
melanism of various insular forms. This is a subject which has been
lately taken up by Dr. Kammerer at the Biologische Versuchsanstalt in
Vienna, and he claims to have produced nigrinos artificially by a very
strong elevation of the temperature, accompanied by extreme dryness. Dr.
Werner[28] has already opposed his own experiments to those of Kammerer,
artificial melanism having been produced by him in _Lacerta oxycephala_
by keeping two very light specimens from Ragusa for a whole summer
in very damp conditions. Neither is Kammerer's theory in accordance
with the distribution of the black lizards, as pointed out by Werner.
Kammerer also finds that those forms which are known to produce melanic
races in a state of nature, lend themselves more readily than the others
to the success of his experiments. But he shows himself misinformed
when he states that the variety called _Lacerta fiumana_ belongs to the
category of those of which black forms are not known. He overlooks the
fact, first pointed out by Scherer in 1904, and which I can confirm,
that the black lizard from Melisello near Lissa in the Adriatic is
unquestionably derived from the lizard from Lissa, which he correctly
regards as not separable from _L. fiumana_...."
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