When, however, he goes on to describe the influences which he regards
as exerted by the habit of freely swimming, I am led to wonder whether
after all in most of these illustrations, the primary distinctions
are not in reality genetic. "Specimens raised in the same aquarium or
in similar aquaria, side by side with all conditions as uniform as it
is possible to make them, seldom fail to furnish striking examples of
broad-headed, short-bodied, and short-tailed types which are habitually
found at the bottom, while others, slender and elongated, are free
swimmers, and maintain themselves in almost as continual suspension
and motion as does a gold fish." Later, again, he writes, "Yet despite
the uniformity of these favourable conditions, the larvae soon began
to split up into two noticeably distinct groups, the one of unusually
compact proportions, the other of uniform intermediate build, such
is most commonly met with." It is to my mind scarcely possible to
resist the inference that, though there may be definite responses
to certain conditions, yet the chief distinctions are genetic, and
that it is these distinctions which confer the power to respond. The
parts respectively played by cause and effect are always difficult to
assign; but when it is stated that "a weak-limbed, long-bodied and
long-tailed animal becomes well nigh perforce an undulatory swimmer,
while the strong-limbed, short-tailed, heavy-bodied specimen, when
these characteristics are rapidly forced upon it, is, under certain
circumstances, just as forcibly induced to become a crawler," we feel
how erroneous any estimates of causation are likely to be.
Public-domain text, read in full here on John Shaqi.
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