In a large series of crosses some seeds died or gave rise to feeble
plants. Of the plants which lived, few gave any seed. The seed, however,
that was obtained from F_{1} plants grew well enough, and the F_{2}
plants proved, as often in such cases, fertile. In these, indeed, no
sign of sterility was noticeable. The experiment is being repeated in
various ways, for, as the genetic behaviour of peas is comparatively
well known, the subject is an exceptionally favourable one for these
investigations.
Such an example shows the confusion produced the moment we attempt to
harmonize conceptions of specific difference with results attained by
experimental methods. It has been usual to regard the field pea (_P.
arvense_) as a species distinct from the edible pea (_P. sativum_).
De Candolle and others regard the field pea as derived from a form
wild in Italy, but the origin of the edible pea is considered to be
unknown. From breeding experiments we find no sterility whatever in the
crosses between the various _arvense_ and _sativum_ types, nor in the
crosses made between them and several other peculiar types from various
countries; whereas this Palestine Pea, which only differs from a small
_arvense_ in what might have been thought trivial characters,[13] either
fails to cross altogether or gives a sterile product, whatever type be
chosen as the other parent.
Examples of this kind have at least the merit that they lead to more
precise delimitations of the problem. We are confronted with two
distinct alternatives.
1. We may apply the term Species promiscuously to all distinct forms.
If we do so it must be clearly understood that we cannot even rule out
the several combinations of "presences and absences" represented by the
various types whether wild or domesticated. For we may feel perfectly
assured that at least all the _arvense_ and all the _sativum_ types yet
subjected to experimental tests are on precisely the same level in this
respect. There is no distinction, logical or physiological, to be drawn
between them. Some contain more factors, and others contain fewer. In
some the re-combinations have been brought about by natural variation or
crossing, while the same consequences in the others have resulted from
man's interference.
2. We may follow the conventions of systematists and distinguish the
outstanding or conspicuous forms such as _arvense_, _quadratum_,
_sativum_ and perhaps a few more as species, and leave the rest
unheeded. If this course is followed it must be clearly understood and
permitted as a piece of pure pragmatism, deliberately adopted for the
convenience of cataloguers and collectors, without regard to any natural
fact or system whatsoever.
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