The three experiments taken together prove, as de Vries says, that the
ovules of _velutina_ are mixed, talls and dwarfs, and that the pollen is
all dwarf. The condition is almost the same as that of the Stocks. It
may be noted also that in the Stocks the egg-cells of the "double" type
are in excess, being approximately 9 to 7 of the "single" type, but de
Vries regards the two types in _velutina_ as probably equal in number.
The figures (169:231) rather suggest some excess of the recessives,
perhaps 9:7, and the point would be worth a further investigation.
As regards _laeta_, by self-fertilisation _no dwarfs were produced_, but
in all other respects it behaved almost exactly like _velutina_. The
ovules are evidently mixed talls and dwarfs, and whether fertilised by
dwarfs or by the pollen of _velutina_, which is already proved to be all
dwarf, the result was a steady 50 per cent. of talls and 50 per cent. of
dwarfs. The pollen of _laeta_ used on dwarfs gives nothing but dwarfs,
and in three series of such experiments 226 dwarfs were produced.
We are thus faced with this difficulty. Since the egg-cells of _laeta_
are evidently mixed, talls and dwarfs, and the pollen used on dwarfs
gives all dwarfs, why does not self-fertilisation give a mixed result,
talls and dwarfs, instead of _all talls_? De Vries regards the result
of self-fertilisation as showing the real nature of the pollen, and
declares it to be all talls, while he represents the behaviour of
the same pollen used on dwarfs by stating that in these combinations
the dwarf character dominates. This does not seem to me a natural
interpretation. I should regard the pollen of _laeta_ as identical
with that of _velutina_, namely dwarf, and I suspect the difficulty
is really created by the behaviour of _laeta_ on self-fertilisation.
Until a proper analysis is made in which the identity of the different
individuals used is recorded, no further discussion is possible.[8]
Other results of a complicated kind involving production of _laeta_ and
_velutina_ together with a third form have been published by de Vries in
his paper on "Triple Hybrids." To these also the same criticism applies.
Some of the observations seem capable of simple factorial representation
and others are conflicting.
Public-domain text, read in full here on John Shaqi.
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