=The Nervous Discharge.=--The word discharge is constantly used, and must
be used in this book, to designate the escape of a current downwards
into muscles or other internal organs. The reader must not understand
the word figuratively. From the point of view of dynamics the passage of
a current out of a motor cell is probably altogether analogous to the
explosion of a gun. The matter of the cell is in a state of internal
tension, which the incoming current resolves, tumbling the molecules
into a more stable equilibrium and liberating an amount of energy which
starts the current of the outgoing fibre. This current is stronger than
that of the incoming fibre. When it reaches the muscle it produces an
analogous disintegration of pent-up molecules and the result is a
stronger effect still. Matteuci found that the work done by a muscle's
contraction was 27,000 times greater than that done by the galvanic
current which stimulated its motor nerve. When a frog's leg-muscle is
made to contract, first directly, by stimulation of its motor nerve, and
second reflexly, by stimulation of a sensory nerve, it is found that the
reflex way requires a stronger current and is more tardy, but that the
contraction is stronger when it does occur. These facts prove that the
cells in the spinal cord through which the reflex takes place offer a
resistance which has first to be overcome, but that a relatively violent
outward current outwards then escapes from them. What is this but an
explosive discharge on a minute scale?
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