In other experiments only a small part of the foot-end was cut from the
animal that served as the graft. The long anterior piece was grafted as
before upon the side of the stock. After the two had united, the graft
was cut in two, leaving a part of the graft attached to the stock. The
part regenerated tentacles, and in two cases subsequently separated from
the stock as in the first experiment. In a third case the graft was
absorbed by the stock as far as the circle of new tentacles, but its
subsequent fate was not determined. In a fourth case the graft did not
regenerate its tentacles, and was completely absorbed into the wall of
the stock. The smaller the piece that is grafted on the stock the
greater the chance that it will be absorbed, and furthermore short,
broad rings are more likely to be absorbed than long, tubular pieces of
the same volume.[80]
Rand’s results show in general that when hydras are grafted together
they regain the typical form in one of two ways,--either by separation
into two individuals, or by the absorption of the smaller into the
larger component. In the former case the result is brought about in the
same way as when the anterior end is partially split in two and the
halves subsequently separate. When the graft is absorbed it is not clear
whether the absorbed piece disappears or, as seems not improbable, forms
a part of the wall of the stock.
It is important to notice the difference between lateral buds and
lateral grafts. The buds separate in the course of four or five days by
constricting at the base, but this never happens in lateral grafts. Rand
has also made some experiments with buds. He cut off the outer oral end
of a bud, and grafted it back upon the stock in a new place. It did not
separate from the stock as does a bud, but by a slow process of division
it was set free in the same way as are lateral grafts. The proximal end
of the bud, which was left attached, developed tentacles at its free
end, constricted at its base, and was set free. The separation was,
however, somewhat delayed.
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