In regard to the origin of the new mesoderm, the evidence is still
insufficient, I think, to show that cells derived from the old muscles
or peritoneum take no part in the formation of the new muscles and
peritoneum; but that the greater part of the new muscles, etc., comes
from the proliferated cells can scarcely be doubted. This latter
discovery loses none of its significance, however, even if it should
prove true that the old muscles, etc., contribute something to the new
part. It is also not entirely disproven that the ventral nerve-cord does
not take a small share in the development of the new cord.
The regeneration of a new tail-end in these same forms appears to take
place in much the same way as the head. The cut-end quickly closes;
later a layer of ectoderm appears over the posterior surface, and the
new part bulges out and becomes dome-shaped. A paired, or in some
species a single, region of proliferation develops from the ectoderm,
that gives rise to the new ventral nerve-cord. Lateral proliferations
of ectoderm produce, according to some writers, the material out of
which the mesoderm of the new tail is formed. Randolph, on the other
hand, has described the new mesoderm as arising from the old, especially
from certain large peritoneal cells that are found throughout the body.
The cut-end of the digestive tract closes, and later new cells develop
at its posterior end. An in-turning of ectoderm, in the form of a pit,
fuses with the posterior end of the digestive tract and establishes
communication with the outside.
[Illustration: FIG 58.--After Hescheler. Regeneration of anterior end of
earthworm. _A._ After four days. _B._ After eleven days. _C._ After
twenty-five days. _D._ After twenty-one days (younger individual).]
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