be given to this result is, I believe, that the new material assumes a
definite structure, or what we may call an organization, and the
subsequent changes are controlled by the kind of structure that is
present; and since this structure has, as a whole, a definite form, we
can state that the form controls the material, although the substitution
of the word “form” for that of “the structure of the new material” may
give the statement an unfortunate, metaphysical appearance.
In order to explain the regeneration of a piece of a plant, Sachs
supposes that two substances are produced by the plant,--one a stem-(or
leaf-) forming substance and the other a root-forming substance. If
either of these substances combines with the protoplasm of any part, a
stem or a root is produced from that part. When a piece of the stem is
cut from a plant, these two substances accumulate, one at the distal end
and the other at the proximal end of the piece, and their presence in
these regions determines that new shoots develop at or near the apex,
and new roots at the base. Sachs tried to show that the direction of the
flow of these two substances is determined by the action of
gravity,--the lighter substance flowing to the higher parts, and the
heavier to the lower parts. We find here reproduced Bonnet’s idea of
specific substances flowing in definite directions; but Sachs goes
farther, and gives an explanation of the cause of the different
directions taken by the two kinds of substances, viz. that it is due to
the action of gravity. Vöchting has shown, as we have seen, after a
thorough examination of the method of development of pieces of plants,
that Sachs’s hypothesis fails to account for the results; and he shows
also that an internal factor, which he calls the polarization, has the
most important influence on the regeneration.
Public-domain text, read in full here on John Shaqi.
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