Vöchting recognizes another sort of influence that determines the
position of new organs on a piece. If a young, growing end of a stem of
_Heterocentron diversifolium_ is suspended by two threads in a
horizontal position, the ends bend upward as a result of the negative
geotropism of the piece. The new roots appear at the base of the piece,
_and also on the convex side of the bent part of the stem_, as shown in
Fig. 34, _B_. The same result can be obtained by forcibly bending a
twig, and then tying the ends together, so that it remains in its bent
position. If a piece of this sort is suspended in a moist atmosphere,
with the bent inner _concave side turned upward_, the roots appear on
the base and at the bend, especially on the _under side_, both from the
nodes and internodes. If now in order to see if gravity takes any part
in the result the next piece is suspended with the outer _convex side_
of the bent part turned _upward_, it is found that many of the pieces
produce roots only at the base, but others produce roots also at the
bent portion of the stem, but they are fewer than in the last
experiment. The roots arise for the most part on the _under side_ of the
arch, and only a few arise from the upper part. It is clear that gravity
is also one of the factors in the result. Leaf-buds arise in these
pieces with the concave side turned _upward_ only near the apex; rarely
one may develop on the lower part of the basal end. In pieces with the
concave side turned _downward_ the leaf-buds arise for the most part at
the apex, but sometimes they appear on the upper part of the basal arm.
The results are due to two factors, gravity and an inner “force” that is
supposed to be the resultant of a growth phenomenon taking place in the
bent portion. Vöchting supposes that a process of growth takes place as
a result of the bending; “the plasma streams to this region, and a new
development takes place here more easily.” Vöchting adds that this view
will not explain the morphological character of the new organs, and that
this must be due to quite other causes. The results may, I venture to
suggest, find a simpler explanation as the result of the bending,
disturbing the tensions of the protoplasm, causing the two arms of the
piece to act as if they had been separated from each other. This idea is
more fully developed in a later chapter.
Public-domain text, read in full here on John Shaqi.
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