In _Hyacinthus
orientalis_, on the other hand, bulblets are produced at the apical
part of a piece of the flowering plant. In this plant the seeds ripen
normally, presumably because of the migration of stuffs toward the
developing seeds. The results in all these cases are due, Goebel thinks,
to the direction of the flow of formative stuffs, and cannot be
explained as connected in any way with the limited growth of the part.
These cases, cited by Goebel, are not in my opinion altogether to the
point; and they fail also to establish convincingly the conclusion that
Goebel draws from them. It may be granted that starch is stored up in
certain parts of the plant, and if these parts are removed the starch
may be stored up in other parts, as Vöchting (’87) has shown; but that
the movement of this starch to the base can account for the lack of
development of the seeds in certain cases seems to me improbable, or, at
least, far from being established by the cases cited. It may be granted
that the presence of starch in a region may act on the organs there
present and determine their fate. Vöchting has shown in the potato that
by removing the tubers the axial buds, especially in the basal leaves,
become tuber-like bodies, but it should not be overlooked that the
tubers themselves are formed from underground stolons, that arise in the
same way as do those in the axils of the leaves. It would be erroneous,
I think, to conclude from these cases of the effect of food stuffs on
certain regions that there are formative stuffs for all the organs of
the plant, and that these stuffs migrate in different directions and
determine the nature of the part. Even the migration of such substances
_in definite directions in the tissue_ is itself in need of explanation,
since it has been made highly probable by Vöchting’s experiments that
this is not produced by agents outside of the plant. Furthermore,
Vöchting has shown that the tendency of starch to accumulate in the
tubers and the formation of the tuber are separate phenomena.
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