The finer tubuli are rarely perfect or fully infiltrated. Even the
coarser canals are not infrequently imperfect, while the laminæ
themselves are sometimes crumpled, crushed, faulted, or penetrated with
veins of chrysotile or of calcite. In some instances the calcareous
laminæ are replaced by dolomite, in which case the canal-systems
are always imperfect or obsolete. The laminæ of the test itself are
also in some cases replaced by serpentine in a flocculent form. At
the opposite extreme are specimens, or portions of specimens, in
which the chambers are obliterated by pressure, or occupied only
with calcite. In such cases the general structure is entirely lost
to view, and scarcely appears in weathering. It can be detected only
by microscopic examination of slices, in parts where the granular
structure or the tubulation of the calcite layers has been preserved.
All palæontologists who have studied silicified fossils in the older
rocks are familiar with such appearances.
[Illustration: Fig. 38.--_Plan of arrangement of Canals in Lamina of
Eozoon._]
It has been alleged by Möbius and others that the canal-systems and
tubes present no organic regularity. This difficulty, however, arises
solely from imperfect specimens or inattention to the necessary results
of slicing any system of ramifying canals. In Eozoon the canals form
ramifying groups in the middle planes of the laminæ, and proceed at
first almost horizontally, dividing into smaller branches, which
ultimately give off brushes of minute tubuli running nearly at right
angles to the surfaces of the lamina, and forming the extremely fine
tubulation which Dr. Carpenter regarded as the proper wall (Figs. 38,
39).
[Illustration: Fig. 39.--_Cross section of minute Tubuli, about 5
microms in diameter (magnified)._]
In my earlier description I did not distinguish this from the
canal-system, with which its tubuli are inwardly continuous. Dr.
Carpenter, however, understood this arrangement, and has represented
it in his figures[30] (see also Fig. 28). It is evident that in a
structure like this a transverse or oblique section will show truncated
portions of the larger tubes apparently intermixed with others much
finer and not continuous with them, except very rarely. Good specimens
and many slices and decalcified portions are necessary to understand
the arrangement This consideration alone, I think, entirely invalidates
the criticisms of Möbius, and renders his large and costly figures of
little value, though his memoir is, as I have elsewhere shown, liable
to other and fatal objections.[31]
[Footnote 30: "Ann. and Mag. Nat. Hist.," ser. 4, xiii., p. 456, figs.
3, 4.]
[Footnote 31: "Museum Memoir," pp. 50 _et seq._]
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