These creatures abound in all parts of the ocean, from the surface
to the greatest depths. The Foraminifera have also existed from
the earliest geological times, and in all the long ages of the
earth's history seem to have retained the same structures and even
ornamentation; so that species from very old geological formations
are often scarcely distinguishable from those now living, and must
have played precisely the same parts in the system of nature. One of
these functions is that of accumulating great thicknesses of calcareous
matter in the sea-bottom.
The manner in which such accumulation takes place we learn from what
is now going on in the ocean, more especially from the result of the
recent deep-sea dredging expeditions. The Foraminifera are vastly
numerous, both near the surface and at the bottom of the sea, and
multiply rapidly; and as successive generations die, their shells
accumulate on the ocean bed, or are swept by currents into banks, and
thus in process of time constitute thick beds of white chalky material,
which may eventually be hardened into limestone. This process is now
depositing a great thickness of white ooze in the bottom of the ocean;
and in times past it has produced such vast thicknesses of calcareous
matter as the chalk and the nummulitic limestone of Europe and the
orbitoidal limestone of America. The chalk, which alone attains a
maximum thickness of 1,000 feet, and, according to Lyell, can be traced
across Europe for 1,100 geographical miles, may be said to be entirely
composed of shells of Foraminifera imbedded in a paste of still more
minute calcareous bodies, the Coccoliths, which are probably products
of marine vegetable life, if not of some animal organism still simpler
than the Foraminifera.
There are, however, some sessile examples of these animals which
attain to larger dimensions than the free and locomotive forms. As an
example of these we may take the _Polytrema_, which forms little hard
red lumps on West Indian corals. Such a creature, beginning life as a
little round spot of protoplasm, almost invisible, and protected with a
little dome of carbonate of lime for the extension of its pseudopods as
it grows in size, adds chamber to chamber in successive tiers till it
assumes an appreciable size, all the chambers communicating with each
other, while the outer ones are perforated with pores for extension of
the pseudopods. In one form (_Carpenteria_) the same end is secured by
leaving an open space in the middle of the conical mass like the crater
of a small volcano. It is with these larger and sessile forms that
we must compare Eozoon, though some of its minute structures rather
resemble those of some smaller types.
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