Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia): Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIIIHaeckel, Ernst
Science
Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia): Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIII
Haeckel, Ernst
Radiolaria
whilst the irregular spheres are due to the mutual dependence of the social
Polyzoa.
118. _The Lattice-Spheres of the Acantharia._--The lattice-shells or
Sphæroid skeletons of the ACANTHARIA are immediately distinguishable from
those of all other Radiolaria by their centrogenous development and the
central union of the radial spines by which they are supported; the only
exception is furnished by the remarkable genus _Cenocapsa_ {lxxix}(Pl. 133,
fig. 11), in which the radial spines are absent, not primitively, however,
but in consequence of degeneration; for the twenty cross-shaped perspinal
pores, originally due to the twenty radial spines, are still present. In
the most nearly allied genera, _Porocapsa_ (Pl. 133, fig. 7) and
_Cannocapsa_ (Pl. 133, fig. 8), the proximal part of the twenty radial
spines is still present, while their distal portion has degenerated; hence
in this case they do not stand in direct communication with the spherical
shell. On the other hand, this primitive connection persists in the genera
_Astrocapsa_ (Pl. 133, figs. 9, 10), and _Sphærocapsa_ (Pl. 135, figs.
6-10). The five genera just mentioned form the peculiar family
Sphærocapsida (pp. 795-802); the spherical shell is in these cases composed
of very numerous small plates disposed like a pavement, each plate or aglet
being perforated by a pore canal; in addition to which there are twenty
larger (perspinal) pores (or twenty cross-shaped groups each of four
aspinal pores) at those important points where primitively the twenty
radial spines penetrate the calymma. This peculiar porous "pavement shell"
has probably been developed (independently of the twenty radial spines)
upon the calymma of the #Acanthonida# (_Acanthonia_, p. 749) by the action
of the sarcodictyum; it has, therefore, quite a different morphological
significance from the spherical lattice-shell of the Dorataspida, which is
composed of tangential apophyses of the twenty Acanthonid spines (pp.
802-847, Pls. 134-138). Each radial spine here forms either two opposite or
four crossed transverse processes, and since their branches spread over the
surface of the spherical calymma and are united suturally at their
extremities, the peculiar lattice-sphere of the Dorataspida arises. This
extensive family is again divided into two subfamilies:--the Diporaspida
(Pls. 137, 138) possess always only two opposite apophyses, and form by the
union of their branches two opposite primary apertures or aspinal meshes.
The Tessaraspida, on the other hand (Pls. 135, 138), have always four
crossed transverse processes, and form by their union four primary aspinal
meshes. From the Diporaspida are probably to be derived the Phractopeltida
(p. 847, Pl. 133, figs. 1-6), the only ACANTHARIA which possess a double
lattice-sphere; their double concentric spherical shell may be compared
with that of the Dyosphærida.
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