Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia): Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIIIHaeckel, Ernst
Science
Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia): Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIII
Haeckel, Ernst
Radiolaria
123. _The Cyrtoid Skeleton._--Cyrtoid skeletons are those lattice-shells
which possess a vertical main axis with two different poles (Monaxonia
allopola); the upper pole is usually termed the apical, the lower the
basal. Such Cyrtoid shells are characteristic of the great majority of the
NASSELLARIA or MONOPYLEA (and especially of the #Cyrtellaria#); they are
also found in a large division of the PHÆODARIA (the #Phæogromia#), and in
some SPUMELLARIA. In general the manifold Cyrtoid shells may be divided
into two large groups, those with one and those with several chambers. The
_monothalamous_ Cyrtoid shells are usually ovoid, conical, cap- or
helmet-shaped; their {lxxxii}internal cavity is simple, without
constrictions or septa. Among the NASSELLARIA they occur in the Monocyrtida
(Pls. 51-54, 98), where they have received the name "Cephalis." A form of
shell, essentially the same, is found amongst the PHÆODARIA in the order
#Phæogromia#, more especially in the Challengerida (Pl. 99), Medusettida
(Pls. 118-120), and Tuscarorida (Pl. 100), many of these latter closely
resembling many Monocyrtida. Such monothalamous Cyrtoid shells occur much
more rarely among the SPUMELLARIA (_e.g._, among the #Prunoidea# in
_Lithapium_, _Lithomespilus_, _Druppatractus_, Pls. 13, 14, &c.).
Polythalamous Cyrtoid shells (Pls. 55-80) occur exclusively in the
NASSELLARIA, and exhibit in this legion an astonishing variety of
structure; they are distinguished from the monothalamous forms by the
development of internal septa, or of annular incomplete diaphragms, which
usually correspond to the external constrictions; their interior is thus
divided into two or more communicating compartments. Among the
polythalamous Cyrtoid shells may be distinguished three principal groups,
the Stichocyrtid, Zygocyrtid, and Polycyrtid. Zygocyrtid shells are
characteristic of the #Spyroidea# (Pls. 84-90), and are distinguished by a
bilobate cephalis (cephalis bilocularis); the median sagittal ring, or a
corresponding constriction, divides the shell into right and left
compartments. Polycyrtid shells (Pl. 96) are peculiar to the #Botryodea#,
and characterised by a multilobate cephalis (cephalis multilocularis).
Stichocyrtid shells are those in which the primary cephalis remains simple,
and new joints are successively added to its basal pole; such shells occur
in the majority of the #Cyrtoidea#. Secondary chambers are sometimes added
in the other two groups (#Botryodea# and #Spyroidea#). When, as often
happens in these polythalamous Cyrtoid shells, two or three distinct joints
follow each other, the first is called the "cephalis," the second the
"thorax," and the third the "abdomen" (Tricyrtida Pls. 64-75).
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