Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia): Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIIIHaeckel, Ernst
Science
Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia): Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIII
Haeckel, Ernst
Radiolaria
125. _The Plectoid Skeleton._--Those forms are distinguished as Plectoid in
which three, four, or more radial siliceous spines proceed from a common
point, which lies excentrically outside the central capsule and at the
basal pole of its vertical allopolar main axis. This peculiar type of
skeletal formation only occurs in the legion NASSELLARIA, and is specially
characteristic of the order #Plectoidea# (p. 898, Pl. 91). But since the
essential elements of this remarkable skeleton also occur in many other
NASSELLARIA, sometimes combined with the Circoid, sometimes with the
Cyrtoid skeleton, it perhaps has a fundamental significance in this legion;
at all events it is possible to derive monophyletically all the other forms
of this legion from it (Hypothesis B, p. 893). The simplest form of the
Plectoid skeleton is a tripod, the three feet of which either lie in a
horizontal plane (_Triplagia_, Pl. 91, fig. 2), or correspond to the three
edges of a low pyramid (_Plagiacantha_). A fourth ray is sometimes added,
which stands vertically upon the summit of the pyramid (_Plagoniscus_,
_Plagiocarpa_, Pl. 91, figs. 4, 5). In other #Plectoidea# three secondary
rays are intercalated between the three primary (Hexaplagida, &c.); seldom
the number is greatly increased (Polyplagida, &c.). The rays are rarely
simple, but usually branched; in the Plagonida (Pl. 91, figs. 2-6) the
branches remain free; in the Plectanida (Pl. 91, figs. 7-13) they are
united to form a loose wicker-work. From such a web a perfect Cyrtoid shell
may arise. Several forms of Plagonida may also be readily confounded with
the isolated triradiate or quadriradiate spicula of many Beloid skeletons
(_Sphærozoum_, _Lampoxanthium_, &c.).
126. _The Spongoid Skeleton._--From the simple lattice-skeleton which the
majority of Radiolaria possess, some of them develop a spongy shell; the
trabeculæ of the lattice-work, situated in one plane in the former, are
developed in the latter in different planes and cross irregularly in all
directions; thus arises a kind of wicker-work of more or less spongy
structure, usually with very thin trabeculæ and irregular meshes. Such
Spongoid shells are most common among the SPUMELLARIA, especially in the
#Sphæroidea# (Spongosphærida, Pl. 18) and #Discoidea# (Spongodiscida, Pls.
41-47), more rarely in the #Prunoidea# and #Larcoidea#. Lattice-work of
similar spongy structure occurs very seldom among the NASSELLARIA, _e.g._,
in some #Plectoidea# (Pl. 91) and #Cyrtoidea# (_Spongocyrtis_,
_Spongopyramis_, _Spongomelissa_, &c., Pl. 56, fig. 10; Pl. 64, figs. 5-10,
&c.). Among the PHÆODARIA spongy skeletons are very rare; they {lxxxiv}are
to be seen in some #Phæosphæria# (_Oroplegma_, Pl. 107, fig. 1;
_Sagoplegma_, Pl. 108, fig. 2; _Auloplegma_, Pl. 111, fig. 8). No Spongoid
skeletons are known among the ACANTHARIA.
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