Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia): Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIIIHaeckel, Ernst
Science
Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia): Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIII
Haeckel, Ernst
Radiolaria
129. _Medullary and Conical Shells._--In all Radiolaria whose skeleton
consists of a double shell or of two concentric lattice-shells united by
radial bars, an inner medullary shell (testa medullaris) and an outer
cortical shell (testa corticalis) may be distinguished (see note A, below).
The medullary shell is usually to be regarded as a primary, the cortical as
a secondary structure. Such double shells occur among the SPUMELLARIA in
the Dyosphærida (Pls. 19, 20), as well as in many #Prunoidea# (Pls. 39,
40), #Discoidea# (Pls. 33, 34), and #Larcoidea# (Pls. 9, 10); among the
ACANTHARIA only in the family Phractopeltida (Pl. 133); among the
NASSELLARIA only in very few #Cyrtoidea# (_e.g._, _Periarachnium_, Pl. 55,
fig. 11), and finally among the PHÆODARIA in the Cannosphærida (Pl. 112) as
well as in part of the Coelodendrida (Pl. 121) and Coelographida (Pls. 127,
128). In most cases (if not always?) the cortical shell arises by the
growth of radial spines from the surface of the medullary shell; these
become united at equal distances from the centre by transverse apophyses,
the surface of the secondary calymma furnishing the basis for their
secretion (§ 85). Nevertheless, it seems that in many #Sphærellaria# the
formation of the whole cortical shell proceeds simultaneously (at a
definite dictyotic period) like that of the primary medullary shell (see
note B). Whilst in the PHÆODARIA, ACANTHARIA, and NASSELLARIA, at most two
concentric shells are formed, in many SPUMELLARIA their number increases
continuously with additional growth; in many #Sphærellaria# it rises to
four, eight, or even more, as well as in many #Discoidea# (if the
concentric, peripherally disposed rings of chambers be regarded as
incomplete flattened shells). In these cases either only the innermost
primary lattice-shell is to be styled "medullary shell," or at most the two
innermost (inner and outer medullary shells), all the others being
cortical.
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