Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia): Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIIIHaeckel, Ernst
Science
Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia): Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIII
Haeckel, Ernst
Radiolaria
134. _Dictyosis of the Phæodaria._--The lattice-structures of the
PHÆODARIA, which consist of a silicate of carbon (§ 102), are on the whole
not developed in such variety as those of the other Radiolaria, but exhibit
several essentially different types of structure, not reducible to a common
primitive type of lattice-work. In one portion of this legion there occurs
an ordinary simple lattice-work (as in SPUMELLARIA and NASSELLARIA), with
solid trabeculæ; of these the Castanellida (Pl. 113) and Concharida (Pls.
123-125) have usually regular or subregular, circular meshes, sometimes
hexagonally framed; the Orosphærida (Pls. 106, 107) large irregular
polygonal meshes with thick trabeculæ, the Sagosphærida (Pl. 108) large
triangular meshes with thin filiform trabeculæ. The Challengerida (Pl. 99)
are characterised by a very delicate regular lattice-work, with minute
hexagonal pores, like a Diatomaceous frustule. The Medusettida (Pls.
118-120) {lxxxix}show a peculiar alveolar structure, numerous small
compartments being enclosed between two parallel plates. In the Circoporida
(Pls. 114-117) and Tuscarorida (Pl. 100) the opaque porcellanous shell has
a peculiar cement structure (§ 104), and the lattice-structure is confined
for the most part to characteristic rings of pores at the base of the
hollow tubes, which arise from the shell. The most peculiar lattice-work,
however, appears in the segmented shell of the Aulosphærida (Pls. 109-111)
and Cannosphærida (Pl. 112). In the former the large meshes of the
lattice-work are usually subregular and triangular, in the latter
polygonal; the trabeculæ are hollow cylinders, filled with jelly, and
containing usually a central axial thread. In each nodal point of the
lattice, in which three or more tangential tubes meet, these are separated
by stellate or astral septa.
135. _Radial Spines of the Skeleton._--The skeleton in the great majority
of Radiolaria is armed with radial spines, which are of great importance in
the development of their general form and of their vital functions. From a
morphological point of view the number, arrangement, and disposition of the
spines is usually the determining factor as regards the general form of the
skeleton. Physiologically they discharge distinct functions, as organs of
protection and support; they act also, like the tentacles of the lower
animals, as prehensile organs, since their points, lateral branches, barbed
hooks, &c. serve to hold fast nutritive materials. In general main-spines
and accessory spines may be distinguished in most Radiolaria; the former
are of pre-eminent importance in determining the figure of the skeleton;
the latter are merely appendicular organs. The main-spines present such
characteristic and important differences in the various legions of
Radiolaria that they must be considered separately.
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