Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia): Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIIIHaeckel, Ernst
Science
Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia): Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIII
Haeckel, Ernst
Radiolaria
all "species bonæ" were "malæ." There may be observed in the manifold
skeletal forms of the Radiolaria, on the one hand, the utmost accuracy of
configuration, and on the other, the greatest variability, and hence a
careful comparative study of them leads to a firm conviction of the gradual
"Transformation of Species," and of the truth of the "Theory of Descent."
157. _Palæontological Development._--The palæontology of the Radiolaria
already offers very considerable material for study; but in consequence of
its incompleteness this is of little value for the study of the phylogeny
of the class. By far the larger portion of the fossil Radiolaria belong to
the Tertiary period; only quite recently have numerous well-preserved
fossil Radiolaria been described from the Mesozoic period, and especially
from the Jura. Of Palæozoic Radiolaria (from the coal measures) only slight
traces are known. Moreover, the fossil Radiolaria hitherto known have been
found only in very circumscribed and widely separated localities. The
majority of all the species belong to the small island of Barbados.
Although our palæontological acquaintance with the Radiolaria must
necessarily be incomplete for this reason, it is still more so since at
least thirty out of the eighty-five families (that is more than a third)
could not possibly leave any fossil remains, either because they possess no
skeleton, or because of its chemical composition.
Of the four legions of the Radiolaria, the ACANTHARIA (on account of the
solubility of their astroid acanthin skeletons) have entirely vanished
and have never been found fossil. Of the PHÆODARIA, whose silicate
skeleton is not as a rule capable of fossilisation, only one section
(Dictyochida) of a single family (Cannorrhaphida) has been observed
fossil. Hence the fossil remains of the Radiolaria belong almost
exclusively to the two legions, SPUMELLARIA and NASSELLARIA, which were
formerly united under the term "Polycystina." Among these, however, the
skeletonless Thalassicollida, Collozoida, and Nassellida could leave no
traces. Hence there only remain fifty-five families of which we might
expect to find fossil siliceous skeletons. Even of these, however,
scarcely the half are certainly known in the fossil condition, whilst of
the remainder nothing certain is known; for example, of the large order
#Larcoidea# (among the SPUMELLARIA) and of the #Stephoidea# (among the
NASSELLARIA) with a few isolated exceptions, no fossils are known. The
great majority of fossil Radiolaria belong to the two NASSELLARIAN orders
#Cyrtoidea# and #Spyroidea# (two relatively very highly developed
groups); next to these follow the orders {cv}#Discoidea# and #Sphæroidea#
among the SPUMELLARIA. From these palæontological facts it is obvious
that our present very incomplete acquaintance with the fossil Radiolaria
is quite insufficient to warrant us in drawing any conclusions from it
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