Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia): Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIIIHaeckel, Ernst
Science
Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia): Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIII
Haeckel, Ernst
Radiolaria
164. _Genealogical Tree of the Sphæroidea._--_Cenosphæra_, the simplest
form of the spherical lattice-shell, may be unhesitatingly regarded as the
common stem-form of all the #Sphæroidea# (pp. 50-284, Pls. 5-30).
_Cenosphæra_ (p. 61, Pl. 12) arose directly from _Actissa_ simply by the
silicification of the spherical exoplasmatic network of the sarcodictyum
around the central capsule, on the surface of the concentric calymma. From
this simple siliceous extracapsular lattice-sphere all other forms of
#Sphæroidea# have arisen, in the main by the manifold combination of two
simple processes, first by the formation of radial spines on the surface of
the lattice-sphere, and second, the addition of concentric spherical
lattice-shells. Both processes may be utilised as the foundation for a
systematic treatment of the #Sphæroidea# (compare pp. 52-58).
If in the #Sphæroidea# the characteristic number and disposition of the
_radial spines_ be regarded as the most important heritable peculiarity
of the different families, then we have the following natural
arrangement:--(1) Liosphærida, without radial spines; (2) Cubosphærida,
with six radial spines (opposite in pairs in three axes perpendicular to
each other); (3) Staurosphærida, with four radial spines (in two axes
crossed at right angles); (4) Stylosphærida, with two opposite radial
spines (in the vertical main axis); and (5) Astrosphærida, with numerous
regularly or irregularly distributed radial spines (eight to twenty or
more). If, on the contrary, more stress be laid upon the number of the
concentric lattice-shells, then we have the following artificial
grouping:--(1) Monosphærida, with one simple lattice-sphere; (2)
Dyosphærida, with two concentric lattice-spheres; (3) Triosphærida, with
three; (4) Tetrasphærida, with four; (5) Polysphærida, with numerous
(five to twenty or more) concentric lattice-shells; (6) Spongosphærida,
with a spongy spherical shell. In general the former arrangement appears
more natural than the latter, since the number of primary radial spines,
which grow out from the primary lattice-sphere, determines their
ground-form from the outset, whatever may be the number of secondarily
added shells. Strictly speaking, according to the view adopted, these
Liosphærida which have several shells, on the outer surface of which
there are no radial spines, ought to be classified according to the
number and arrangement of their internal radial connecting beams and
distributed among the other families. The practical application of this
correct principle meets, however, with great difficulties. Also in many
cases the phylogenetic relations of the different #Sphæroidea# are more
complicated than would appear from both these classificatory principles.
In general their phylogeny will quite correspond with their ontogeny,
since from the innermost first formed {cix}lattice-shell (primary
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